Opuntia abjecta

Opuntia abyssi

Opuntia acanthocarpa

Opuntia acanthocarpa ssp. acanthocarpa

C.B. Wolf, Occasional Papers [Rancho Santa Ana Botanic Garden] 1: 75, 1938

L.D. Benson, The cacti of Arizona 20, 34, f. 1. 4, 1. 5, 1969

Opuntia acanthocarpa v. major

Opuntia acanthocarpa v. ramosa

Opuntia acanthocarpa v. thornberi

Opuntia acaulis

Opuntia aciculata

Opuntia aciculata v. orbiculata

Opuntia acracantha

Opuntia aequatorialis hyb.

Opuntia affinis

Opuntia aggeria

Opuntia agglomerata

Opuntia airampo

Opuntia alamosensis

Opuntia albata

Salm-Dyck, Hort. Dyck. 361.,

Salm-Dyck, Cact. Hort. Dyck. 67, 237. 1850, 1849

Scheinvar, Revista Jard. Bot. Nac. Univ. Habana 20: 167 (-169). 1999

K. Schumann, Gesamtb. Kakt. Nachtr. I. 152.,

Backeberg, Kaktus-ABC 127, 411, 1935

Backeberg, Cactaceae (Backeberg) 6: 3617., 1962

Hildmann ex Hirscht, Monatsschr. Kakteenk., 10. 48, in obs., pro syn., 1900

Mieckley ex Lindemuth & Dams, Monatsschr. Kakteenk., 14. 178, in obs., nomen., 1904

F.A.C. Weber, Bulletin du Museum d'Histoire Naturelle 1: 321, 1895

J. Iliff, Cactaceae Consensus Initiatives 2: 9, 1997

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 4: 256, 1923

(Spegazzini) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

(Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

Karwinski ex Salm-Dyck, Cact. Hort. Dyck. ed. 2. 68,

Karwinski ex Foerst., Ruempl. Handb. Kakteenk. ( ?) 938, sphalm., 1885

Cárdenas, Lilloa 23: 23, f. 3, 1950

D. Griffiths, Illustrated Studies in the Genus Opuntia. -- 2. 83-84, t. 5, t. 2, f. 2, t. 12, upper f., t. 13, f. 9, 1909

K. Schumann ex Dams, Monatsschr. Kakteenk., 13. 95, in obs., 1903

Gill. ex Pfeiffer, Enum. Diagn. Cact. 146,

D. Griffiths, Annual Report of the Missouri Botanical Garden 21: 165166, t. 19, 20, upper fig., 1910

Griffiths, Bull. Torr. Bot. Cl., 46. 205., 1919

Link ex Tenore, Atti 7 Adun. Sci. Ital. 884, 1846

Small, Journal of the New York Botanical Garden 20: 29, 1919

Tenore, Fl. Neap. Prod. App. 5: 15, 1826

Spegazzini, Bulletin du Museum d'Histoire Naturelle 10: 391, 1904

(Spegazzini) R. Kiesling, Candollea, 53 (2): 475: 1998

(Spegazzini) R. Kiesling, Candollea, 53 (2): 475: 1998

(Spegazzini) R. Kiesling, Candollea, 53 (2): 476: 1998

D. Griffiths, Bulletin of the Torrey Botanical Club 43: 92, 1916

Pfeiffer, Enumeratio Diagnostica Cactearum 145, 1837

Lemaire, Cact. Gen. Sp. Nov. 72., 1839

Lemaire, Cact. Gen. Sp. Nov. 72., 1839

Lemaire, Cact. Gen. Sp. Nov. 72., 1839

Meinsh., C. Koch & Fint. Wochenschr. 1. 30., 1858

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 292, 1857

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 425, 1896

D.J.Pinkava, Madrono; Journal of the California Botanical Society 23 (5): 293, f. 5, 1976

Britton & Rose, Brooklyn botanic garden memoirs 1: 73, 1918

Lemaire, Cactearum aliquot novarum 34, 1838

(F. Ritter) R. Crook & R. Mottram, Bradleya; Yearbook of the British Cactus and Succulent Society 13: 100, 1995

F.A.C. Weber, Bulletin du Museum d'Histoire Naturelle 4: 165, 1898

Steud., Nom. ed. 2. 2. 220.,

Engelmann, Memoir of a Tour to Northern Mexico 90, 1848

Dams, Monatsschrift fur Kakteenkunde 14: 3, 1904

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 309, 1857

(Griffiths) G. D. Rowley, Natl. Cact. Succ. J. 13: 25., 1958

Cárdenas, Cactus and Succulent Journal [U. S. ] 28: 113, f. 77-78, 1956

(F. Ritter) D.R. Hunt, Cactaceae Consensus Initiatives 4: 5, 1997

Spegazzini, Anal. Mus. Buenos Aires, 11. 520.,

Engelmann, Proc. Amer. acad. 3: 301, 1856

Grisebach, Symbolae ad Floram Argentinam. Zweite . 140, 1874

D. Griffiths, Illustrated Studies in the Genus Opuntia. -- 2. 93-94, t. 10, t. 2, f. 8, t. 13, f. 8, 1909

Engelmann ex Hirscht, Monatsschrift fur Kakteenkunde 8: 115, 1898

Backeberg, Cactus (Paris) 38: 250, 1953

Backeberg, Descr. Cact. Nov. 9., 1956

Backeberg, Cactus (Paris) 8 (36): 181, 1953

(Pfeiffer) D.R. Hunt, Bradleya; Yearbook of the British Cactus and Succulent Society 5: 93, 1987

(Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13. 5, 1958

K. Schumann, Monatsschrift fur Kakteenkunde 153, 1899

R. Philippi, Florula Atacamensis seu enumeriation . 24, 1860

D. Griffiths, Annual Report of the Missouri Botanical Garden 21: 172-173, t. 26. lower fig., 1910

Small, Manual of the Southeastern Flora 905, 1506, 1933

(Backeberg ex F. Ritter) R. Crook & R. Mottram, Bradleya; Yearbook of the British Cactus and Succulent Society 13: 106, 1995

Rose, Smithsonian Miscellaneous Collections 50: 518, 1908

Spegazzini, Anales de Sociedad Cientifica Argentina 99: 91, 1925

Werdermann & Backeberg, Neue Kakteen 63, 1931

(Rauh & Backeberg) Krainz, Die Kakteen, 1970

(Rauh & Backeberg) Krainz, Die Kakteen, 1970

(Rauh & Backeberg) Krainz, Die Kakteen, 1970

Pfeiffer, Allgemeine Gartenzeitung 8: 282, 1840

F.A.C. Weber, Bull. Mus. HIst. Nat. Par., 392., 1904

Lindl., Edwards's Botanical Register 19: pl. 1606, 1833

Baxter, Cactus and Succulent Journal [U. S. ] 489, 1933

(Brack & Heil) D.J. Pinkava & B.D. Parfitt, Sida 13 (2): 128, 1988

F.A.C. Weber, Dictionaire d'Horticulture 896, 1898

Small, Flora of the Southeastern United States . 816, 1335, 1903

Rose, Contributions from the U. S. National Herbarium 12 (7): 291, t. 24, f. 33, 1909

G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 203, f. 254-255, 1919

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 210-211, f. 271, 1919

J.E. Madsen, Flora of Ecuador 35: 47, 1989

Weber ex Hirscht, Monatsschr. Kakteenk., 8. 175, in obs., pro syn., 1898

Rose, Contributions from the U. S. National Herbarium 13 (9): 309, t. 64, 1911

K. Brandegee ex J.A. Purpus, Monatsschr. Kakteenk., 10. 97, in obs., 1900

K. Brandegee ex J. A. Purpus, Monatsschr. Kakteenk. 10: 110, 120., 1900

(Daston) Rowley, Nat. Cact. & Succ. Journ. 13 (1): 5, 1958

Rafin., Atlantic Journal 146, 1832

Engelmann & J. M. Bigelow, Proc. Amer. Acad. 3: 298, 1856

(E.M. Baxter) Munz, Aliso 4 (1): 94, 1958

F.A. Walton 

(Griffiths) Munz ex L.D. Benson, Nat. Cacti Calif. 128, 1969

Fobe, Monatsschrift fur Kakteenkunde 16: 46, 1906

S.L. Welsh & Neese, Great Basin Naturalist 43 (4): 700, 1983 [1984]

(Griffiths) W.T. Marshall, Cactaceae 65, 1941

(Clover & Jotter) L. D. Benson, Cacti Ariz. 2nd. ed. 43., 1950

Parish, Bulletin of the Torrey Botanical Club 19: 92, 1892

(J.M. Coulter) Toumey 

W. Earle, Saguaroland Bulletin 34: 15, 1980

K. Schumann, Gesamtb. Kakt. 1. 722.,

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 111, f. 137-139, 1919

H. Sánchez-Mejorada R., Cactaceas y suculentas mexicanas 17 (2): 47, f. 40, 1972

D. Griffiths, Annual Report of the Missouri Botanical Garden 22: 25, 1912

F.A.C. Weber, Gardener's chronicle, ser. 3 35: 34, 1904

Engelmann ex Parish, Bulletin of the Torrey Botanical Club 19: 92, 1892

(Colla) A.E. Hoffmann J., Cactaceas en la flora silvestre de Chile: 244: 1989

R. Philippi, Linnaea 33: 83, 1864

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 307, 1856

Fosberg, Bulletin of the Southern California Academy of Sciences 32: 121, 1933

Pittier, Journal of the Washington Academy of Sciences 26: 42, 1936

Backeberg, Cactus, Paris, No. 75, 32; Backeb. Cactac. Handb. Kakteenk. 6. 3607 (1962). 1962

Rose, Contributions from the U. S. National Herbarium 12: 402, 1909

(Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 155, t. 26, f. 3, 1919

Salm-Dyck, Allgemeine Gartenzeitung 13: 388, 1845

Backeberg, Kaktus-ABC 128, 411, 1935

Spegazzini, Contr. Flora Tandil 18, 1901

(Kunth) F.A.C. Weber, Enumeratio Diagnostica Cactearum,

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 103-104, f. 126, 1919

(F. Ritter) R. Crook & R. Mottram, Bradleya, 13: 115: 1995

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 302, 1856

D. Griffiths, Proceedings of the Biological Society of Washington 27: 25, 1914

(Griffiths) Wiggins & C.B. Wolf, Ill. Fl. Pacific States 3: 148., 1951

K. Schumann, Kuntze, Rev. Gen. 3. 2. 107,

(Daston) Rowley, Nat. Cact. & Succ. Journ. 13 (1): 5, 1958

G.D. Rowley, National Cactus and Succulent Journal 13: 4, 1958

(J.M. Coulter) K. Brandegee, Erythea 5: 121, 1897

K. Schumann, Gesamtbeschreibung der Kakteen 1: 652, 1898

(Willd.) Haworth, Supplementum Plantarum Succulentarum . 79, 1819

(Britton & Rose) P.J. Braun & E. Esteves Pereira, Succulenta, 74 (3): 132: 1995

(C.T. Rizzini & A. Mattos) P.J. Braun & E. Esteves Pereira, Succulenta, 74 (3): 132: 1995

Baxter, Anales del Instituto de Biologia de la Universidad Nacional de Mexico 4: 149, 1933

H. E. Gates, Cactus and Succulent Journal [U. S. ] 10: 98, 19938

J.G. Ortega, Mex. Forest 4 (4): 68, 1928

Spegazzini, Physis. Revista de la Sociedad Argentina de Ciencias Naturales 8: 239, 1925

Spegazzini, Revista Argent. Bot. 1: 202., 1926

(F. Ritter) C. Schlindwein, Bradleya, 13: 29: 1995

Britton & Rose, Cactaceae, 1. 150., 1919

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 304-305, 1856

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 70-71, t. 14, f. 1, 1919

Griffiths & Hare, New Mexico Agricultural Experiment Station: Bulletin 60: 47, 1906

D. Griffiths, Illustrated Studies in the Genus Opuntia. -- 2. 86-87, t. 6, upper f., t. 2, f. 9, 1909

Griffiths, Proc. Biol. Soc. Wash., 29. 13., 1916

Rafin., Bulletin Botanique [Geneve] 216, 1830

Forbes ex Baxt., Loud. Hort. Brit. Suppl. 3. 599, nomen.,

Griffiths, Bull. Torr. Bot. Cl, 43. 524., 1916

Wherry, Journal of the Washington Academy of Sciences 16: 12, f. 1, 1926

Engelmann, Notes of a Military Reconnoissance 2: 158, 1848

(J.M. Coulter) D.J. Pinkava, Haseltonia 4: 103, 1996

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 453, 1896

Lemaire, Cact. Gen. Sp. Nov. 73,

Espinosa, Revista Chilena de Historia Natural 37: 126, 1933

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 293, 1856 [1857]

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 99, f. 120, 1919

M.A. Baker & D.J. Pinkava, Cact. Succ. J. (U. S. A.), 71 (6): 320, 1999

F.A.C. Weber, Dictionaire d'Horticulture 895, 1898

D. Griffiths, Illustrated Studies in the Genus Opuntia. -- 2. 90-91, t. 6, lower f., t. 2, f. 6, t. 13, f. 2, 12, 1909

Hort. Monac. ex Pfeiffer, Enum. Diagn. Cact. 159.,

Salm-Dyck, Cact. Hort. Dyck. 68, 241. 1850, 1849

Spegazzini, Anal. Mus. Buenos Aires, 11. 518.,

Lynch, Gardener's chronicle, ser. 3 33: 98, 1903

Arechavaleta, Anales del Museo Nacional de Montevideo 5: 278, 1905

Salm-Dyck, Cacteae in Horto Dyckensi Cultae [ed. 1849] 238, 1850

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 259-260, t. 21, 1908

K. Schumann, Monatsschrift fur Kakteenkunde 9: 150, 1899

F.A.C. Weber, Dictionaire d'Horticulture 895, 1898

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 49, f. 59, 1919

Usteri, Guia Bot. Praca Rep. e Jard. Luz, 27., 1919

Forbes, Hort. Tour Germ. 160, nomen.,

R. Philippi, ex K. Schum. Gesamtb. Kakt., Nachtr., 152, pro syn., 1903

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 261-262, t. 24, 1908

Werdermann, Notizbl. Bot. Gart. Berlin, 12. 223., 1934

Hort. Berol. ex Pfeiffer, Enum. Diagn. Cact. 166.,

Backeberg, Backeb. & F. M. Knuth, Kaktus-ABC, 128, 411., 1935

F.A.C. Weber, Dictionaire d'Horticulture 897, 1898

Spath, Cat. -7. 156; cf. Kew Bull. 1907, App. 3. 74, hybr. ?, 1906

Rafin., Ser. Bull. Bot. 216, 1830

Griffiths, Proc. Biol. Soc. Wash., 27. 25., 1914

Britton & Rose, Contributions from the U. S. National Herbarium 16 (7): 241, t. 72, 1913

Spegazzini, Anal. Mus. Buenos Aires, 11. 519., 1905

F.A.C. Weber, Bulletin du Museum d'Histoire Naturelle 10: 388, 1904

Clokey, Madrono; Journal of the California Botanical Society 7: 71, 1943

Griffiths, Bull. Torrey Bot. Club 46: 199. 1919,

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 264-265, t. 23, lower fig., 1908

Weber., Bull. Mus. Hist. nat. Paris. 1. 320., 1895

(Cárdenas) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

(Cárdenas) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

Rose, Contributions from the U. S. National Herbarium 12 (7): 291-292, 1909

(Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13: 5, 1958

K. Koch, Hortus Dendrologicus 279, no. 6, 1853

(Anthony) D.J. Ferguson, Cactus and Succulent Journal [U. S. ] 58 (3): 124, 1986

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 291, 1856

(Weber) D.J. Ferguson, Cact. Succ. J. (U. S. A.), 60 (4): 159: 1988

Griffiths & Hare, New Mexico Agricultural Experiment Station: Bulletin 60: 64, 1906

F.A.C. Weber, Bulletin du Museum d'Histoire Naturelle 1: 320, 1895

Cárdenas, Lilloa 23: 20, f. 2, 1950

Wiggins, Contributions from the Dudley Herbarium 3: 285, 1943

Engelmann, Contributions from the U. S. National Herbarium 3: 445, 1896

Pfeiffer, Enumeratio Diagnostica Cactearum 173, 1837

Schelle, Handb. Kakteenkult. [Schelle] 43., 1907

(A. Cast.) G. D. Rowley, Natl. Cact. Succ. J. 13: 25., 1958

Engelmann, Memoir of a Tour to Northern Mexico 95, 1848

Engelmann ex J.M. Coulter, Contributions from the U. S. National Herbarium 3: 444, 1896

Hort. ex Pfeiffer, Enum. Diagn. Cact. 161,

Steud., Nom. ed. 2. 2. 220,

Cárdenas, Revista de Agricultura [Cochabamba] 9: 20, 1955

Karst., Deutsche Fl. 888 (-83), in obs., 1880

(L.) Mill., The Gardeners Dictionary: eighth edition Opuntia no. 6, 1768

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 263-264, t. 26, 1908

Gill. ex Pfeiffer, Enum. Diag. Cact. 155,

(F. Ritter) P.J. Braun & E. Esteves Pereira, Succulenta (Netherlands) 74 (3): 132, 1995

F. A. C. Weber in Bois, Dict. Hort. 2: 895., 1898

(F. Ritter) D.R. Hunt, Cactaceae Consensus Initiatives 4: 5, 1997

A. Castellanos, Lilloa 27: 81, f. 1, 1955

D. Griffiths, Bulletin of the Torrey Botanical Club 43: 523, 1916

(J.M. Coulter) Britton & Rose, Smithsonian Miscellaneous Collections 50: 519, 1908

(Salisb.) J.F. Macbride, Contributions from the Gray Herbarium of Harvard University 65: 41, 1922

(Griffiths) D. Weniger, Cacti Southwest 207, 1970

(Small) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 124, 1969

(Small) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 125, 1969

(Engelmann) D. Weniger, Cacti Southwest 207, 1971

D. Weniger, Cacti S. W. 202, 1970

(Engelmann) L.D. Benson, Cact. Ariz. 50, 1950

(Engelmann) L. D. Benson, Proceedings of the California Academy of Sciences, Series 4, 25: 250, 1944

(Engelmann & J. M. Bigelow) D. D. Weniger, Cacti Southwest 211, 1970

D. Griffiths, Proceedings of the Biological Society of Washington 27: 28, 1914

D. Griffiths, Illustrated Studies in the Genus Opuntia. -- 2. 88-89, t. 8, t. 2, f. 4, 7, f. 13, f. 5, 1909

(F. Ritter) P.J. Braun & E. Esteves Pereira, Succulenta (Netherlands) 74 (3): 133, 1995

(F. Ritter) G.D. Rowley, National Cactus and Succulent Journal 13: 25, 1958

Mackensen, Bulletin of the Torrey Botanical Club 39: 291, 1912

(Small) Backeberg, Neue Kakteen, 1931

Spegazzini, Anales del Museo Nacional de Buenos Aires 11: 513,

K. Schumann ex D. Vaupel, Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 50 (2-3, Beibl. 111): 28-29, 1913

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

Salm-Dyck, Cacteae in Horto Dyckensi Cultae [ed. 1844] 43, 1844

Salm-Dyck, Cact. Hort. Dyck. 72. 1850, 1849

Miq., Nederl. Kruidk. Arch. 4. 838., 1859

Britton & Rose, Smithsonian Miscellaneous Collections 50: 532, 1908

Munz, Man. S. Calif. Bot. 327., 1935

(Fosberg) Munz, A Manual of Southern California Botany 327, 1935

Haworth, Supplementum Plantarum Succulentarum . 81, 1819

Pfeiffer, Enumeratio Diagnostica Cactearum 153, 1837

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

(F. Ritter) D.R. Hunt, Cactaceae Consensus Initiatives 4: 5, 1997

D. Griffiths, Proceedings of the Biological Society of Washington 29: 11, 1916

Salm-Dyck, Enumeratio Diagnostica Cactearum 157, 1837

Pfeiffer, Enumeratio Diagnostica Cactearum 157, 1837

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

(Rauh & Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

Salm-Dyck, Cact. Hort. Dyck. ed. I. 49; et ed. 2. 78,

Hort ex Pfeiffer, Enum. Diagn. Cacl. 167,

D. Griffiths, Bulletin of the Torrey Botanical Club 43: 528, 1916

Britton & Rose, Torreya 12: 14, 1912

Griffiths, Bull. Torr. Bot. Cl., 43. 524., 1916

(Griffiths & Hare) Britton & Rose, Smithsonian Miscellaneous Collections 50: 529, 1908

Hort. ex Schelle, Handb. Kakteenk. 46, in syn., 1907

Small, Manual of the Southeastern Flora 907, 1506, 1933

(L.) Mill., The Gardeners Dictionary: eighth edition Opuntia no. 7, 1768

Backeberg, Descr. Cact. Nov. 10., 1956

Haworth, Revis. Pl. Succ. 71., 1821

Griffiths (pro sp.), Bulletin of the Torrey Botanical Club 43: 88, 1916

D. Griffiths, Annual Report of the Missouri Botanical Garden 22: 30, t. 9-10, 1910

Griffiths, Bull. Torrey Bot. Club 46: 196. 1919,

Griffiths, Rep. (Annual) Missouri Bot. Gard. 22: 30, pl. 9, 10., 1912

(Engelmann & J. M. Bigelow) Rose, Contributions from the U. S. National Herbarium 13 (9): 309, 1911

(Cárdenas) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

(Lam.) DC., Prodromus Systematis Naturalis Regni Vegetabilis 3: 471, 1828

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 295, 1856

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 296, 1857

D. Vaupel, Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 50 (2-3, Beibl. 111): 29, 1913

F.A.C. Weber, Bulletin du Museum d'Histoire Naturelle 1904: 388, 1904

Hensl., Magazine of Zoology and Botany 1: 466, 1837

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 305, 1856

Rose, Contributions from the U. S. National Herbarium 13 (9): 309, pl. 65, 1911

De Candolle, Memoires du Museum d'Histoire Naturelle 17: 118, 1828

(Willd.) Haworth, Supplementum Plantarum Succulentarum . 71, 1821

Salm-Dyck, Hortus Dyckensis ou Catalogue des Plantes . 361, 1834

Eichlam, Monatsschr. Kakteenk. 21: 116., 1911

E. Paniagua 

Lemaire, Cact. Gen. Sp. Nov. 68,

Salm-Dyck, Hortus Dyckensis ou Catalogue des Plantes . 361, 1834

Weber, Monatsschr. Kakteenk., 14. 166, in obs., nomen; Vaupel, Bluhende Kakteen, 37. t. 148 (1913), descr. Vaupel, Bluhende Kakteen, 1904, 14. 166, in obs., nomen; 37. t. 148 (1913), descr., 1904

Rose, Contributions from the U. S. National Herbarium 13 (9): 310, 1911

D. Griffiths, Bulletin of the Torrey Botanical Club 43: 84, 1916

D. Griffiths, Rep. (Annual) Missouri Bot. Gard. 22: 29, pl. 8., 1912

Forbes, Hott. Tour Germ. 160, nomen,

Hort. Monac. ex Foerst., Handb. Cact. ed. I. 508, nomen.,

(Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13. 4, 1958

B.E. Ralston & R.A. Hilsenbeck, Madrono; Journal of the California Botanical Society 39 (4): 281-284, f. 1, 1992

F. A. C. Weber, Dict. Hort. 2: 897., 1898

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 216, f. 279, 1919

Wendl. ex Foerst., Handb. Cact. ed. I. 508, nomen.,

Rose, Smithsonian Miscellaneous Collections 50: 517, 1908

D. Griffiths, Monatsschrift fur Kakteenkunde 2: 132, 1913

Lemaire, Cact. Aliq. Nov. Desc. 36.,

(Lemaire) Schelle, Handb. Kakteenkult. [Schelle] 45., 1907

Spegazzini, Anales Mus. Nac. Buenos Aires (ser. 3) 4: 511., 1905

Spegazzini, Anales Mus. Nac. Buenos Aires (ser. 3) 4: 511., 1905

Spegazzini, Anales Mus. Nac. Buenos Aires (ser. 3) 4: 511., 1905

Hort. ex Forbes, Hort. Tour Germ. 157.,

Hott. ex Pfeiffer, Enum. Diagn. Cact. 159,

A. Web., Bois, Dict. Hortic. 898,

A. Weber, Bulletin du Museum d'Histoire Naturelle 4: 166, 1898

D. Griffiths, Illustrated Studies in the Genus Opuntia. -- 2. 82-83, t. 4, lower f., t. 2, f. 10, t. 13, t. 7, 1909

(Ker Gawl.) Haworth, Supplementum Plantarum Succulentarum . 79, 1819

Salm-Dyck, Hortus Dyckensis ou Catalogue des Plantes . 185, 1834

Salm-Dyck, Cacteae in Horto Dyckensi Cultae [ed. 1849] 67, 1849 [1850]

L. Scheinvar, Feddes Repert., 95 (5-6): 277, 1984

H. Bravo Hollis, Cactaceas y suculentas mexicanas 9: 55, 1969

C. F. Forster, Hamburger Garten- und Blumenzeitung 17: 167, 1861

(Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 25., 1958

Hort. ex A. Berger, Hort. Mortol. 232, nomen., 1912

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 266-267, t. 27, upper fig., 1908

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 218, f. 282, 1919

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 155-156, f. 195, 1919

Griffiths, Bull. Torrey Bot. Club 46: 197, pl. 9. 1919,

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 225, f. 301, 1919

Hort. ex Pfeiffer, Enum. Diagn. Caut. 167,

(F. Ritter) D.R. Hunt, Cactaceae Consensus Initiatives 4: 5, 1997

Britton & Rose ex Urb., Symb. Antill. 8. 466, nomen; Britton & Rose, Cactaceae, 4. 259 (1923), in obs.,, 1920

R. Graham, Botanist 5: t. 246, 1846

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 291, 1857

A. Berger, Kakteen 58, 1929

Britton & Rose, Smithsonian Miscellaneous Collections 50 (4): 518, 1908

(Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13. 5, 1958

(Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

Foerst. ex Haage, Verz. Cact. 29,

Lemaire, Cact. Aliq. Nuv. Desc. 85,

Small, The Cactaceae; descriptions and illustrations of plants of the cactus family 4: 260, 1923

(F. Ritter) A.E. Hoffmann J., Cactaceas en la flora silvestre de Chile: 248: 1989

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 305, 1856

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 305, 1856

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 446, 1896

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 446, 1896

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 446, 1896

L.D. Benson, Cact. Calif. 93, 1969

J.T. Howell, Proceedings of the California Academy of Sciences, Series 4, 21: 49, 1933

E.Y. Dawson, Cactus and Succulent Journal [U. S. ] 34: 104, 1962

J.T. Howell, Proceedings of the California Academy of Sciences, Series 4, 21: 51, 1933

E.Y. Dawson, Cactus and Succulent Journal [U. S. ] 34: 103, 1962

E.Y. Dawson, Cactus and Succulent Journal [U. S. ] 34: 104, 1962

J. T. Howell, Proc. Calif. Acad. Sci. 21: 51., 1933

(Howell) E.F. Anderson & Walk., Madrono; Journal of the California Botanical Society 20: 256, 1970

V.E. Grant & K.A. Grant, Botanical Gazette 140 (2): 205, 1979

Griffiths, Bull. Torrey Bot. Club 46: 195. 1919,

A. Berger, Contributions from the U. S. National Herbarium 13 (9): 310, pl. 66, 1911

Werdermann, Repertorium Specierum Novarum Regni Vegetabilis 29: 225, 1931

Link & Otto ex Salm-Dyck, Hortus Dyckensis ou Catalogue des Plantes . 861, 1834

F. A. C. Weber, Bull. Mus. Hist. Nat. (Paris) 10: 392., 1904

E. Walther, J. Cact. Succ. Soc. Amer. 1: 204, figs. 2, 4., 1930

E. Walther, J. Cact. Succ. Soc. Amer. 1: 204, figs. 2, 4., 1930

Miller, The Gardeners Dictionary: eighth edition Opuntia no. 4, 1768

E. Sanchez-Mejorada & Villasenor, Acta Botanica Mexicana 27: 1, 1994

Miquel, Nederlandsch Kruidkundig Archief. Verslangen en Mededelingen der Nederlandsche Botanische Vereeniging 4: 337, 1858

D.A. Griffiths, Annual Report of the Missouri Botanical Garden 21: 170-171, t. 25, 1910

Haworth, Suppl. Pl. Succ. 81,

Salm-Dyck, Cact. Hort. Dyck. 242. 1850, 1849

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 304, 1856

Salm-Dyck, Cacteae in Horto Dyckensi Cultae [ed. 1849] 67, 235, 1849 [1850]

(Engelm.) Schelle, Geological Survey of California, Botany 1: 248, 1876

(Griffiths) D. Weniger, Cacti S. W. 178, 1969

(Griffiths) D. Weniger, Cacti S. W. 175, 1970

(Griffiths & Hare) D. Weniger, Cacti S. W. 177, 1970

Griffiths & Hare, New Mexico Agricultural Experiment Station: Bulletin 60: 44, t. 2, 1906

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 291, 1857

(Griffiths) A. Nelson, Transactions of the Academy of Science 12: 124, 1919

(Engelmann) J.M. Coulter ex K. Schumann, Gesamtbeschreibung der Kakteen 725, 1898

(Peebles) L.D. Benson, Cacti Ariz. 58, 1950

(L.D. Benson) B.D. Parfitt & D.J. Pinkava, Madrono; Journal of the California Botanical Society 35 (4): 348, 1988

(Griffiths) B.D. Parfitt & D.J. Pinkava, Madrono; Journal of the California Botanical Society 35 (4): 348, 1988

(Engelmann) B.D. Parfitt & D.J. Pinkava, Madrono; Journal of the California Botanical Society 35 (4): 346-347, 1988

(Griffiths) B.D. Parfitt & D.J. Pinkava, Madrono; Journal of the California Botanical Society 35 (4): 347, 1988

(Griffiths) Fosberg, Bulletin of the Southern California Academy of Sciences 33: 100, 1924

Engelmann 

(Griffiths) D. Weniger, cacti S. W. 180, 1970

(Griffiths) D. Weniger, Cacti S. W. 174, 1970

Fosberg, Bull. S. Calif. Acad. Sci. 30: 59., 1931

Griffiths, Proc. Biol. Soc. Wash., 29. 11., 1916

Hort. ex K. Schumann, Monatsschr. Kakteenk., 11. 181, in obs., nomen., 1901

Backeberg, Kaktus-ABC 128, 411, 1935

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 301, 1856

(E. M. Baxter) S.L. Welsh, Great Basin Naturalist 46 (2): 255, 1986

(Griffiths) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 124, 1969

(Engelmann & J. M. Bigelow) L.D. Benson, Cact. Ariz. 46, 1940

Dunkle, Bulletin of the Southern California Academy of Sciences 34: 3, 1935

(K. Schumann) L. D. Benson, Proc. Calif. Acad. Sci. 14th ser. 25: 249., 1944

(F.A.C. Weber) Parish, A Flora of California 2: 542, 1936

(Engelmann) L.D. Benson, The cacti of Arizona 20, 1969

(K. Schumann) L.D. Benson 

Lemaire, Handbuch der Kakteenkunde 492, 1846

P.J. Braun, Cactus and Succulent Journal [U. S. ] 62 (4): 165, 1990

A. Berger, Hortus Mortolensis, 232, 410., 1912

H. Sánchez-Mejorada R., Cactus and Succulent Journal [U. S. ] 17 (3): 67, 1972

D. Griffiths, Proceedings of the Biological Society of Washington 29: 14, 1916

Salm-Dyck ex Pfeiffer, Enum. Diagn. Cact. 147.,

De Candolle, Memoires du Museum d'Histoire Naturelle 17: 118, 1828

DC., Mem. Mus. Hist. Nat. nat. 17: 118. Revue fam. Cact. 118. 1829, 1828

DC., Mem. Mus. Hist. Nat. nat. 17: 118. Revue fam. Cact. 118. 1829, 1828

Lemaire, Cact. Gen. Sp. Nov. 67., 1839

Salm-Dyck, Cact. Hort. Dyck. 48. 1845, 1844

Lemaire ex Labour., Cact. 492,

Ekman & Werdermann, Repertorium Specierum Novarum Regni Vegetabilis 29: 228, 1931

Britton & Rose, Contributions from the U. S. National Herbarium 23: 1676, 1926

Rose ex J.G. Ortega, Fl. Indig. Sinaloa, 1929

(Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13. 5, 1958

Haworth, Suppl. Pl. Succ. 82.,

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 267-268, 1908

Sencke ex Haage, Verz. Cact. 29 (Quid ?).

Haworth 

A. Berger, Monatsschrift fur Kakteenkunde 22: 181, 1912

(L.) Mill., The Gardeners Dictionary: eighth edition Opuntia n. 2, 1768

(Haworth) Spegazzini, Anales del Museo Nacional de Buenos Aires 4: 512, 1905

(F.A.C. Weber) Spegazzini, Anales del Museo Nacional de Buenos Aires 3 (4): 512, 1905

Backeberg, Descr. Cact. Nov. 10., 1956

(Ten.) A. Berger, Handb. Kakteenkult. [Schelle] 51., 1912

Schelle, Handb. Kakteenkult. [Schelle] 51., 1907

Makino, Journal of Japanese Botany 7 (4): 6, f. 1, 1931

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 294, 1856

Peebles, Cactus and Succulent Journal [U. S. ] 29: 67, 1937

Lemaire, Cactearum Genera Nova Speciesque Novae 61, 1839

Hort. Berol. ex Foerst., Handb. Cact. ed. I. 508, nomen.,

Hort. ex Pfeiffer, Enum. Diagu. Cact. 161,

D. Griffiths, Bulletin of the Torrey Botanical Club 43: 87, 1916

(Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13: 5, 1958

Salm-Dyck, Allgemeine Gartenzeitung 13: 388, 1845

(Rauh & Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

(Marn.-Lap.) Rowlev, Rep. Pl. Succ., 22 (Regnum Veg., 87): 11, 1973

(Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

F. A. C. Weber in Bois, Dict. Hort. 2: 897., 1898

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

Lemaire, Cact. Gen. Sp. Nov. 68,

Salm-Dyck, Prodromus Systematis Naturalis Regni Vegetabilis 3: 471, 1828

F.A. Walton, Cact. Journ. 1. 105., 1898

C. B. Wolf (pro sp.), Occasional Papers [Rancho Santa Ana Botanic Garden] 1: 79, 1938

(Nutt.) Haworth, Synopsis plantarum succulentarum . 182, 1819

(Engelmann & J. M. Bigelow) W.A. Weber, Phytologia 51 (6): 374, 1982

L.H. Bailey, The Standard Cyclopedia of Horticulture 4: 2363, 1916

Wiegand & Backeberg, Descriptiones Cactacearum Novarum 10, 1956

Engelmann, Boston Journal of Natural History 5: 245, 1845

Backeberg, Descriptiones Cactacearum Novarum 10, 1956

L.H. Bailey, The Standard Cyclopedia of Horticulture 4: 2363, 1916

(F. Ritter) G. Navarro, Lazaroa 17: 66, 1996

Gibbes, Proceedings of the Elliott Society of Natural History 1: 273, 1859

Engelmann, Boston Journal of Natural History 6: 208, 1850

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 309, 1857

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 309, 1857

Engelmann, Bot. Calif. 1. 250,

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 306, 1857

(Fosberg) L.D. Benson, Cact. Calif. 106, 1969

(Schott ex Engelmann) J.M. Coulter, Contributions from the U. S. National Herbarium 3: 449, 1896

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 262-263, t. 25, 1908

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

Salm-Dyck, Enumeratio Diagnostica Cactearum 164, 1837

Salm-Dyck, Cact. Hort. Dyck. 65, 234. 1850, 1849

Salm-Dyck, Enumeratio Diagnostica Cactearum 164, 1837

Wendland & Pfeiffer, Enumeratio Diagnostica Cactearum 170, 1837

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 297, 1857

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 271, t. 23, upper left fig., 1908

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 297, 1856 [1857]

Hemsl., Magazine of Zoology and Botany 1: 467, 1837

Opuntia galapageia subvar. inermis

Backeberg, Kakteenlexikon 317., 1966

(J. T. Howell) Backeberg, Die Cactaceae 1: 561, 1958

(J. T. Howell) Backeberg, Die Cactaceae 1: 562, 1958

(K. Schumann ex Robins) Backeberg, Die Cactaceae 1: 562, 1958

(Stewart) Backeberg, Die Cactaceae 1: 561, 1958

E.Y. Dawson, Cactus and Succulent Journal [U. S. ] 37: 141, 1965

(F. A. C. Weber) Backeberg, Die Cactaceae 1: 561, 1958

E.F. Anderson & Walk., Madrono; Journal of the California Botanical Society 20: 256, 1970

(J. T. Howell) Backeberg, Die Cactaceae 1: 562, 1958

(J. T. Howell) Backeberg, Die Cactaceae 1: 562, 1958

De Smet ex Miq., Nederl. Kruidk. Arch. 4. 337., 1859

(F. Ritter) D.R. Hunt, Cactaceae Consensus Initiatives 4: 5, 1997

(C.B. Wolf) J. Rebman & D.J. Pinkava, Haseltonia 4: 103, 1996

R. Philippi, Anal. Univ. Chil. 85. 492., 1894

A. Castellanos, Kakteenk., 172., 1934

Hort. Berol. ex Pfeiffer, Enum. Diagn. Cact. 102,

D. Griffiths, Annual Report of the Missouri Botanical Garden 87-88, t. 7, t. 2, f. 5, t. 13, f. 6, 1909

D. Griffiths, Annual Report of the Missouri Botanical Garden 22: 35, t. 9, 16, 17, 1912

Hort. Berol. ex Pfeiffer, Enum. Diagn. Cact. 158,

Forbes, Hort. Tour Germ. 158, 1837

Salm-Dyck, Hortus Dyckensis ou Catalogue des Plantes . 362, 1834

Wendl., Cat. Hort. Herrnh.; ex Pfeiff. Diagn. Cact. 162-, 1835

Salm-Dyck, Cact. Hort. Dyck. 66. 1850, 1849

Haworth, Philosophical magazine, or annals of chemistry, 7: 111, 1830

(R. Philippi) Castell., Lilloa 23: 12., 1950

(Lemaire) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

(Salm-Dyck) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

(Spegazzini) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

(Spegazzini) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

(Spegazzini) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

K. Schumann, Gesamtb. Kakt. 654. 1898, 1898

D. Griffiths, Annual Report of the Missouri Botanical Garden 21: 167-168, t. 21, 22, lower fig., 1910

D. Griffiths, Monatsschrift fur Kakteenkunde 23: 134, 1913

F.A.C. Weber, Bulletin de la Societe Nationale d'Acclimatation de France 49: 83, 1902

(Griffiths & Hare) L.D. Benson, Cact. Ariz. 65, 1950

(Wiegand & Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13 (1): 5, 1958

Lemaire, Cact. Gen. et Sp. Nov. 68.,

Pfeiffer, Enumeratio Diagnostica Cactearum 172, 1837

Salm-Dyck, Cact. Hort. Dyck. 73, 251. 1850, 1849

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 304, 1856

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 295, 1856 [1857]

Pfeiffer, Enumeratio Diagnostica Cactearum 155, 1837

G. Phil., Linnaea 30: 211, 1859-1860

Schelle, Handb. Kakteenkult. [Schelle] 46., 1907

Britton & Rose, Smithsonian Miscellaneous Collections 50: 523, 1908

Engelmann ex Coulter, Contrib. U. S. Nat. Herb. 3. 431, in syn., 1896

D. Griffiths, Annual Report of the Missouri Botanical Garden 22: 26, 1912

Mackensen, Bulletin of the Torrey Botanical Club 39: 291, 1912

F.A.C. Weber, Bulletin du Museum d'Histoire Naturelle 1904: 391, 1904

Gurke, Monatsschr. Kakt. xvlii. 180., 1908

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 218, f. 285, 1919

D.R. Hunt, Cactaceae Consensus Initiatives 4: 5, 1997

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 266, 1908

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 265-266, 1908

F.A.C. Weber, Dictionaire d'Horticulture 893, 1898

A. Berger, Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 36: 456, 1905

Britton, Smithson. Misc. Coll. I. 513., 1908

Spegazzini, Anales de Sociedad Cientifica Argentina 99: 97, 1925

(H. E. Gates ex Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13. 25, 1958

A. Weber ex A. Berger, Gard. Chron., I. 34., 1904

Britton & Rose, Johnston & Tryon, Rep. Prickly Pear Comm. 97., 1914

Arp, Sida 10: 207-210, 1984

G.K. Arp, Sida, 10 (3): 207, as 'heacockae', 1984

Scheinvar, Anales del Instituto de Biologia de la Universidad Nacional Autonoma de Mexico 45: 75, 1974

Matuda, Cactaceas y suculentas mexicanas 1 (2): 23-24, f. 14, 1955

K. Schumann ex B.L. Rob., Proceedings of the American Academy of Arts and Sciences 38: 180, 1902

K. Schumann, Gesamtbeschreibung der Kakteen 1: 690, [1898]1899

De Candolle, Mem. Hist. Nat. Paris 17: 69, t. 16, 1828

Rol.-Goss., Bulletin du Museum National d'Histoire Naturelle 15: 507, 1909

Kupper, Monatsschrift fur Kakteenkunde: 212, 1930

R. Philippi, Anales del Museo Nacional. Santiago de Chile 1891: 28, 1891

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 93, f. 108, 1919

Grisebach, Goett. Abh. 24. 140., 1879

Fric, Mollers Deutsch. GartnerZeit., 42. 377., 1927

(Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

Griffiths, Bull. Torrey Bot. Club 46: 198. 1919,

J.G. Ortega, Bol. Pro-Cult. Reg. 1: 13, 1930

Forbes, Hort. Tour Germ. 159, nomen.,

H. Bravo Hollis, Anales del Instituto de Biologia de la Universidad Nacional Autonoma de Mexico, Serie Biologia 1: 89, 1930

Standley, Field Museum of Natural History, Botanical Series 9 (4): 316, f. 8, 1940

Hort. ex Pfeiffer, Enum. Diagn. Cact. 159,

Salm-Dyck, Prodromus Systematis Naturalis Regni Vegetabilis 3: 472, 1828

Heinrich, Backeb. Descr. Cact. Nov. 3. 10., 1963

(Krainz & Graser) Rowley, Nat. Cact. & Succ. Journ. 13. 5, 1958

J.A. Purpus, Mitteilungen der Deutschen Denrologischen Gesellschaft 1925: 61, 1925

H. Bravo Hollis, Anales del Instituto de Biologia de la Universidad Nacional de Mexico 25: 484, 1954

Hester, Cact. Succ. J. (Los Angeles) 15: 191, fig. 94. 1943,

(Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13. 4, 1958

(Raf.) Raf., Medical Flora 2: 247, 1830

(Engelmann) R. Crook & R. Mottram, Bradleya, 16: 135: 1998

(Small) L.D. Benson, Cactus and Succulent Journal [U. S. ] 48: 59, 1976

(Small) Dress, Baileya 19: 165, 1975

(Raf.) Raf. 

(Engelm.) A. Heller, Catalogue of North American Plants North of Mexico (ed. 2) 8, 1900

A. Heller, Catalogue of North American Plants North of Mexico (ed. 2) 8, 1900

A. Heller, Cat. N. Amer. Pl. (ed. 2) 8., 1900

Haworth, Synopsis plantarum succulentarum . 189, 1812

D. Griffiths, Bulletin of the Torrey Botanical Club 43: 381, 1916

Werdermann in Backeberg, Neue Kakteen 64, 1931

Spegazzini, Anal. Mus. Buenos Aires, 11. 509.,

F.A.C. Weber, Dictionaire d'Horticulture 894, 1898

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 299, 1856

Backeberg, Cactaceae (Backeberg) 1: 609., 1958

(L. D. Benson) Backeberg, Cactaceae (Backeberg) 1: 610., 1958

(F.A.C. Weber) Backeberg, Die Cactaceae 1: 610, 1958

Grisebach, Cact. Pl. Cub. 117, 1866

(F. Ritter) Iliff, Cactaceae Consensus Initiatives 2: 8, 1997

D. Griffiths, Monatsschrift fur Kakteenkunde 23: 138, 1913

D. Vaupel, Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 50 (2-3, Beibl. 111): 30, 1913

(Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 5., 1958

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 99, f. 121, 1919

(Haworth) DC., Prodromus Systematis Naturalis Regni Vegetabilis 3: 471, 1828

M. S. Anthony, American Midland Naturalist 55 (1): 225, 1956

(Griffiths) H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 119, 1972

(Rose) H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 119, 1972

Salm-Dyck, Cact. Hort. Dyck. 249. 1850, 1849

Salm-Dyck, Cact. Hort. Dyck. 249. 1850, 1849

Small, The Cactaceae; descriptions and illustrations of plants of the cactus family 4: 257-258, f. 235, 1923

A. Berger, Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 36: 453, 1905

(F. Ritter) R. Crook & R. Mottram, Bradleya, 17: 113: 1999

K. Schumann, Flora Brasiliensis 4 (2): 306, 1890

(F. Ritter) P.J. Braun & E. Esteves Pereira, Cact. Succ. J. (U. S. A.), 61 (6): 272: 1989

Backeberg, Descr. Cact. Nov. 10., 1956

(Schott ex Griffiths) D.R. Hunt, Cactaceae Consensus Initiatives 4: 5, 1997

D. Griffiths, Annual Report of the Missouri Botanical Garden 22: 27, 1912

(DC.) DC., Prodromus Systematis Naturalis Regni Vegetabilis 3: 437, 1828

(F. Ritter) Iliff, Cactaceae Consensus Initiatives 2: 8, 1997

Stewart, Proceedings of the California Academy of Sciences, Series 4, 1: 113, 1911

Salm-Dyck, Hortus Dyckensis ou Catalogue des Plantes . 364, 1834

Salm-Dyck, Cacteae in Horto Dyckensi Cultae [ed. 1849] 69 (1849), 1850

D. Griffiths, Proceedings of the Biological Society of Washington 29: 10, 1916

T. S. Brandeg., Proceedings of the California Academy of Sciences, Series 2, 2: 163, 1889

Otto ex Foerst., Handb. Cact. ed. I. 505,

Cárdenas, Cactus, No. 34, 127; cf. Cact. & Succ. Journ. Amer. 26. 84 (1954). 1952

Mart. ex Pfeiffer, Enum. Diagn. Cact. 154,

Tenore ex Pfeiffer, Enum. Diagn. Cact. 149,

Griffiths, Bull. Torr. Bot. Cl., 43. 529., 1916

H. Bravo Hollis, Cactaceas y suculentas mexicanas 17: 115, 1972

Britton & Harris, Torreya 11: 130, 1911

F.A.C. Weber, Cact. Util. Mexico 120, 1928

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 197, f. 243-244, 1919

Haage ex Foerst., Handb. Cact. ed. I. 508, nomen.,

Salm-Dyck, Cacteae in Horto Dyckensi Cultae [ed. 1849] 239, 1849[1850]

V.E. Grant & K.A. Grant, Evolution, 25 (1): 155., 1971

Britton & Small, Journal of the New York Botanical Garden 20: 31, 1919

Spegazzini, Anal. Mus. Buenos Aires, 11. 516., 1905

De Candolle, Memoires du Museum d'Histoire Naturelle 17: 118, 1828

Fisch., Tax. Rel. 41, 1962

Salm-Dyck, Cact. Hort. Dyck. 73, 250. 1850, 1849

(Toumey) W. T. Marshall, Des. Bot. Gard. Ariz. Sci. Bull. 1: 17, 1950

Werdermann & Backeberg, Neue Kakteen 64, 1931

Werdermann & Backeberg, Neue Kakteen 64, 1931

Backeberg, Cact. & Succ. Journ. Amer. 3. 132, in obs., 1932

Rose, Smithsonian Miscellaneous Collections 50: 505, 1908

(E. M. Baxter) Peebles, Leaflets of Western Botany 5 (12): 192, 1949

Console, Gesamtbeschreibung der Kakteen 717, 1898

Backeberg, Cactus, Paris, No. 75, 32; Backeb. Cactac. Handb. Kakteenk. 6. 3617 (1962), 1962

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 419, 1896

(Griffiths) Peebles, Cact. Succ. J. (Los Angeles) 9: 68., 1937

K. Schumann, Gesamtbeschreibung der Kakteen 151, 1903

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 6, 1958

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1); 5, 1958

Baxter ex H. Bravo Hollis, Anales del Instituto de Biologia de la Universidad Nacional de Mexico 3: 181, 1932

Fric., Ceskoslov. Zahradnickych Listu [Kakt. Sukk. ], 1924, 122, nomen; cf. Gray Herb. Card Cat., Issue 114., 1924

Haworth, Syn. Pl. Succ. 192.,

Parment., Cacteae in Horto Dyckensi Cultae [ed. 1849] 65, 1849[1850]

F.A.C. Weber, Contributions from the U. S. National Herbarium 3: 423, 1897

Pfeiffer, Enumeratio Diagnostica Cactearum 160, 1837

Small, Journal of the New York Botanical Garden 20: 26, 1919

D. Griffiths, Bulletin of the Torrey Botanical Club 43: 85, 1916

Hort. ex A. Berger, Hort. Mortol. 233, 1912

Console ex Bois, Dict. Hort. 894., 1898

Werdermann, Notizblatt des Botanischen Gartens und Museums zu Berlin-Dahlem 10 (97): 752, 1929

Haage & Schmidt 

F.A.C. Weber, Bulletin du Museum d'Histoire Naturelle 10: 393, 1904

Mackensen, Bulletin of the Torrey Botanical Club 38: 141, 1911

De Candolle, Memoires du Museum d'Histoire Naturelle 17: 118, 1828

(Engelmann) S. Watson, Bibliographical Index to North American Botany 1: 407, 1878

(Engelmann) A. Berger, Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 36: 459, 1905

(Engelmann) S. Watson, Bibliographical Index to North American Botany 1: 407, 1878

A. Berger, Kakteen 59, 1929

(Gonzalez Ortega) H. Bravo Hollis, Cact. Suc. Mex. 17: 119., 1972

(J.G. Ortega) H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 119, 1972

A. Berger, Kakteen 59, 1929

A. Berger, Kakteen (Berger) 59., 1929

A. Berger, Kakteen 59, 1929

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 456, 1896

Salm-Dyck, Hortus Dyckensis ou Catalogue des Plantes . 362, 1834

Salm-Dyck, Cacteae in Horto Dyckensi Cultae [ed. 1844] 47, 1844[1845]

Salm-Dyck, Handbuch der Kakteenkunde 497, 1846

Link & Otto, Verh. Ver. Befoerd. Gartenb. 6. 484, nomen; Pfeiff. Enum. Diagn. Cact. 167., 1830

R. Philippi, Anales del Museo Nacional. Santiago de Chile 1891: 27, 1891

Forbes, Hort. Tour Germ. 160, nomen,

Wendland, Enumeratio Diagnostica Cactearum 167, 1837

De Candolle, Memoires du Museum d'Histoire Naturelle 17: 119, 1828

F.A.C. Weber, Gesamtbeschreibung der Kakteen 157, 1903

B. Trujillo & M. Ponce, Ernstia 58-60: 1, 1990

(F. Ritter) P.J. Braun & E. Esteves Pereira, Succulenta (Netherlands) 74 (3): 133, 1995

Engelmann, Boston Journal of Natural History 6: 207, 1850

(Griffiths) H. Bravo Hollis, Cactaceas y suculentas mexicanas 19 (2): 47, 1974

M. S. Anthony, American Midland Naturalist 55: 252, f. 26, 1956

(Griffiths & Hare) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 125, 1969

(Engelmann & J. M. Bigelow) J.M. Coulter, Contributions from the U. S. National Herbarium 3: 422, 1896

(Engelmann) J.M. Coulter, Contributions from the U. S. National Herbarium 3 (7): 421, 1896

(Griff.) K. Hammer, Kulterpflanze, 24: 268, 1976

L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 125, 1969

(Griffiths) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 125, 1969

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 422, 1897

(Griffiths) L. Scheinvar, Phytologia, 49 (4): 320: 1981

J. M. Coulter, Contr. U. S. Natl. Herb. 3: 421., 1896

(Griffiths) J. Elizondo E. & Wehbe, Cact. Suc. Mex. 32: 17., 1987

(Griffiths) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 125, 1969

J.P. Rebman, Cactus and Succulent Journal [U. S. ] 69 (2): 67, 1997

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 270-271, t. 27, lower fig., 1908

(Engelmann) Britton & Rose, Smithsonian Miscellaneous Collections 50: 529, 1908

L.D. Benson & Walk., Ann. Missouri Bot. Gard. 52: 269., 1965

(L.D. Benson) L.D. Benson, Annals of the Missouri Botanical Garden 52: 270, 1965

(Fosberg) L.D. Benson & Walk., Ann. Missouri Bot. Gard. 52: 270., 1965

(Coulter) L.D. Benson & Walk., Ann. Missouri Bot. Gard. 52: 268., 1965

D.R. Hunt, Cactaceae Consensus Initiatives 4: 6, 1997

Rose, Contributions from the U. S. National Herbarium 12 (7): 292, t. 25, f. 34, 1909

Clover & Jotter, Bulletin of the Torrey Botanical Club 68: 418, f. 6, 1941

D. Griffiths, Bulletin of the Torrey Botanical Club 43: 525, 1916

C. Z. Nelson, Trans. Illinois State Acad. Sci. 12: 120. Galesburg Rep. Reg. July 20, 1915, 1919

Haworth, Philosophical magazine, or annals of chemistry, ?: 109, 1830

Backeberg, Cactaceae (Backeberg) 6: 3609., 1962

Backeberg, Cactus (Paris) no. 38: 250., 1953

(Salm-Dyck) Backeberg, Cactus (Paris) no. 38: 250., 1953

Backeberg, Cactus (Paris) no. 38: 250., 1953

Backeberg, Cactus (Paris) no. 38: 250., 1953

Peebles, Cactus and Succulent Journal [U. S. ] 9: 109, 1938

D. Griffiths, Annual Report of the Missouri Botanical Garden 21: 169, t. 23, 1910

(E. M. Baxter) Backeberg, Cactaceae (Backeberg) 1: 585., 1958

Britton (pro sp.), Bulletin of the New York Botanical Garden 4: 141, 1906

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 269-270, 1908

Hort., ex Wien. Ill. Gart. -Zeit. 14. 146, 1889

Hort. ex Borg, Cacti, 88, in syn., 1937

(Rose) J.C. Solomon, Novon 9 (3): 422, 1999

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 221, f. 292-293, 1919

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 4: 261, f. 238, 1923

Rauh & Backeberg, Descriptiones Cactacearum Novarum 1956: 10, 1957

Rose, Smithsonian Miscellaneous Collections 50: 516, 1908

Rose, Contributions from the U. S. National Herbarium 13 (9): 310, t. 67, 1911

Gibbes, Proceedings of the Elliott Society of Natural History 1: 273, 1859

Gibbes, Proc. Elliott Soc. Nat. Hist. 1. 273., 1858

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 268-269, t. 28, upper fig., 1908

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 292, 1856 [1857]

S. Brack & K.D. Heil, Cactus and Succulent Journal [U. S. ] 60: 17, 1988

L.D. Benson, Cact. Ariz. ed. 2 64, 1950

M. S. Anthony, American Midland Naturalist 55: 244, f. 21, 1956

Engelmann, Boston Journal of Natural History 6: 206, 1850

R.E.M. Hernández Valencia, Acta Cient. Potosina, 10 (2): 155, 1988

(Salm-Dyck) L.D. Benson, The cacti of Arizona 20, 1969

C. F. Forster, Hamb. Gartenz. 17. 166., 1861

Hort. Belg. ex Salm-Dyck, Cact. Hort. Dyck. ed. I. 46,

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 268, 1908

D. Griffiths, Proceedings of the Biological Society of Washington 27: 23, 1914

D. Griffiths, Proceedings of the Biological Society of Washington 29 (3): 9-10, 1916

Small, Manual of the Southeastern Flora 906, 1933

Noronha, Verhandelingen van het bataviaasch genootschap van kunsten en wetenschappen 5 (4): 22, 1790

Griffiths, Bull. Torr. Bot. Cl., 46. 201., 1919

Remy, Flora Chilena 3: 29, 1847

Arechavaleta, Anal. Mus. Montevid. 5. 286.,

(W. Rausch) W. Rauh, Kakteen an ihren Standorten: 93, without basionym ref., 1979

Schott ex Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 308, 1856 [1857]

Engelmann, Syn. Cact. U. S. 52., 1856

(Backeberg) G.D. Rowley, Nat. Cact. & Succ. Journ. 13. 6, 1958

W.E. Parsons, Desert Plant Life 8: 10, 1936

(J.M. Coulter) Baxter, Cactus and Succulent Journal [U. S. ] 2: 388, 1931

Rafin., Atlantic Journal 146, 1832

Backeberg, Cactus (Paris) 8 (36): 181, 1953

(L.D. Benson) B.D. Parfitt, Systematic Botany 5 (4): 416, 1980 [1981]

L. Scheinvar, Phytologia, 49 (4): 324, 1981

Miller, The Gardeners Dictionary: eighth edition Opuntia no. 5, 1768

J.G. Ortega, Fl. Ind. Sinaloa, 1929

Haworth, Suppl. Pl. Succ. 82,

Salm-Dyck, Hort. Dyck. 363., 1834

(Pfeiffer) Berger, Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 36: 453, 1905

Salm-Dyck, Cact. Hort. Dyck. 68. 1850, 1849

D. Griffiths, Illustrated Studies in the Genus Opuntia. -- 2. 91, t. 2, f. 3, t. 13, f. 10, 1909

Griffiths, Bull. Torr. Bot. Cl., 43. 530., 1916

Rose, Smithson. Misc. Coll. l. 529., 1908

Arechavaleta, Anales Museo Nacional Montevideo, ser. 2 1: 42, 1905

Rose, Contributions from the U. S. National Herbarium 10: 126, 1906

J.T. Howell, Proceedings of the California Academy of Sciences, Series 4, 21: 46, 1933

J. T. Howell, Proc. Calif. Acad. Sci. 21: 47., 1933

J. Lundh, Madrono; Journal of the California Botanical Society 20: 254, 1970

J.T. Howell, Proceedings of the California Academy of Sciences, Series 4, 21: 48, 1933

VS 187 

Svenson, American Journal of Botany 33: 471, 1946

Rafin., Bulletin Botanique [Geneve] 216, 1830

(Engelmann & J. M. Bigelow) J.M. Coulter, Contributions from the U. S. National Herbarium 3: 430, 1896

(Engelmann) J.M. Coulter, Contributions from the U. S. National Herbarium 3: 429, 1896

Engelmann ex J.M. Coulter, Contributions from the U. S. National Herbarium 3: 431, 1896

(Engelmann) J.M. Coulter, Contributions from the U. S. National Herbarium 3: 430, 1896

(Engelmann & J. M. Bigelow) J.M. Coulter, Contributions from the U. S. National Herbarium 3: 429, 1896

Engelmann ex J.M. Coulter, Contributions from the U. S. National Herbarium 3 (7): 431, 1896

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 429, 1896

J.M. Coulter, Contributions from the U. S. National Herbarium 19: 446, 1915

(Engelmann & J. M. Bigelow) J.M. Coulter, Contributions from the U. S. National Herbarium 3: 430, 1896

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 431, 1896

Picol. ex Salm-Dyck, Cact. Hort. Dyck. ed. I. 46,

Pittier, Journ. Wash. Acad. Sc., 26. 41., 1936

Hort. ex Pfeiffer, Enum. Diagn. Cact. 150, 160,

D. Griffiths, Monatsschrift fur Kakteenkunde 23: 130, 1913

K. Schumann, Gesamtbeschreibung der Kakteen 714, 1898

(Lehmann) Pfeiffer, Enumeratio Diagnostica Cactearum 154, 1837

Salm-Dyck, Bot. Gart. 186, 1834

Backeberg, Descriptiones Cactacearum Novarum 10, 1956

Fobe, Monatsschrift fur Kakteenkunde 235, 1931

Schelle, Kakteen 60, 1926

Salm-Dyck, Cacteae in Horto Dyckensi Cultae [ed. 1849] 241, 1849

Salm-Dyck ex Schelle, Kakteen 60, 1926

Borg, Cacti 83, 1951

(Engelmann) K. Schumann, Gesamtbeschr. Kakt. 706., 1898

F.A.C. Weber, Dictionaire d'Horticulture 896, 1898

Hort. ex Pfeiffer, Enum. Diagn. Cact. 173,

K. Schumann, Bluhende Kakteen 4: pl. 44, 1903

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 104, f. 127, 1919

Britton, Smithsonian Miscellaneous Collections 50: 513, 1908

(Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13. 6, 1958

C. Muell., Walp. Ann. 5. 50, sphalm.,

(Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13: 6, 1958

(Backeberg) A. Cast., Lilloa 23: 12, 1950

Monville, L'Horticulteur Universel 1: 218, 1839

G.D. Rowley, Natl. Cact. Succ. J. 13: 4., 1958

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

De Candolle, Prodromus Systematis Naturalis Regni Vegetabilis 3: 472, 1828

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 300, 1856

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 300, 1856 [1857]

Salm-Dyck, Cact. Hort. Dyck. 46. 1845, 1844

Späth 

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 300, 1856 [1857]

Engelmann, Proc. Amer. Acad. Arts 3: 300., 1856

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 300, 1856

Späth 

Hort 

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 300, 1856 [1857]

(Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

A. Berger, Kakteen 57, 1929

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 293, 1856 [1857]

T. S. Brandeg., Proceedings of the California Academy of Sciences, Series 2, 2 (2): 164, 1889

(Spegazzini) Werdermann, Feddes Repertorium 30, 59, 1932

Spegazzini, Anales de Sociedad Cientifica Argentina 96: 63, 1923

(Willd.) Haworth, Suppl. Pl. Succ. 81., 1819

(F. Ritter) P.J. Braun & E. Esteves Pereira, Succulenta, 74 (3): 133: 1995

Salm-Dyck, Cact. Hort. Dyck. 66, 234. 1850, 1849

Salm-Dyck, Cact. Hort. Dyck. 66, 235. 1850, 1849

(L.) Haworth ex Steud., Nomenclator Botanicus. Editio secunda 2: 221, 1841

(Areces) R. Govaerts, World Checklist Seed Plants, 3 (1): 21: 1999

Sencke, Haage, Verz. Cact. 29 (Quid ?).

Spegazzini, Anal. Mus. Buenos Aires, 11. 515; et ex Arech. in Anal. Mus. Montevid. 5. 282., 1905

R. Philippi, Linnaea, 33. (-65) 82., 1864

K. Schumann, Kuntze, Rev. Gen. 3. 2. 107, nomen.,

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 47, t. 6, f. 1, 1919

(Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13. 6, 1958

Backeberg, Cactus, Paris, No. 75, 32; Backeb. Cactac. Handb. Kakteenk. 6. 3614 (1962). 1962

Clokey, Madrono; Journal of the California Botanical Society 7: 69-70, pl. 4, f. a, 1943

C.B. Wolf, Occasional Papers [Rancho Santa Ana Botanic Garden] 1: 79, 1938

F.A.C. Weber, Dictionaire d'Horticulture 894, 1898

Vis., Flora Dalmatica 3: 143, 1852

Britton, Bulletin of the New York Botanical Garden 3: 446, 1905

(Areces) R. Govaerts, World Checklist Seed Plants, 3 (1): 21: 1999

H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 115, 1972

Rose, Smithsonian Miscellaneous Collections 50: 516, 1908

D. Griffiths, Monatsschrift fur Kakteenkunde 23: 133, 1913

D. Griffiths, Annual Report of the Missouri Botanical Garden 19: 260-261, t. 23, upper fig., 1908

(Backeberg) G.D. Rowley, Nat. Cact. & Succ. Journ. 13. 4, 1958

H. Bravo Hollis, Cactaceas y suculentas mexicanas 19 (1): 20, 1974

Y. Ito, The Cactaceae: 85, nom. nov., without full repl. syn. ref.: 1981

Borg, Cacti, 67., 1937

L.D. Benson, Cacti Arizona, ed. 2, 48; cf. Gray Herb. Card Cat., 1950

(Haworth) Haworth, Handb. Caot. ed. I. 496,

K. Schumann, Gesamtbeschreibung der Kakteen 1: 695, 1898 [1899]

D. Griffiths, Annual Report of the Missouri Botanical Garden 21: 169-170, t. 24, 1910

Small, Manual of the Southeastern Flora 906, 1933

(Backeberg & H. Jacobsen) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

Karwinski ex Salm-Dyck, Cact. Hort. Dyck. ed. 2. 68,

(Backeberg) G.D. Rowley, Repertorium Plantarum Succulentarum 22: 11, 1973

Backeberg, Cactus, Paris, No. 75, 32; Backeb. Cactac. Handb. Kakteenk. 6. 3614 (1962), 1962

Wendland, Enumeratio Diagnostica Cactearum 161, 1837

Griffiths, Bull. Torr. Bot. Cl., 46. 202., 1919

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 291, 1856

(Britton & Rose) Parish, Man. Fl. Pl. Calif. 657., 1925

(Engelmann) Parish, A Manual of the Flowering Plants of California . 657, 1925

(Griffiths) Munz, Aliso 4 (1): 94, 1958

(Fosberg) Munz, Aliso 4 (1): 94, 1958

(J.M. Coulter) Munz, Aliso 4 (1): 94, 1958

Small, The Cactaceae; descriptions and illustrations of plants of the cactus family 4: 262-263, 1923

DC. in DC., Pl. Hist. Succ. 2: 138, fig., 1804

Salm-Dyck, Cacteae in Horto Dyckensi Cultae [ed. 1849] 241, 1849[1850]

Engelmann ex J.M. Coulter, Contributions from the U. S. National Herbarium 3: 431, 1896

(L.) Karst., Deutsche Flora. Pharmaceutisch-medicinische Botanik. 888, 1882

Salm-Dyck ex Pfeiffer, Enumeratio Diagnostica Cactearum 156: 1837

Philbrick, Cact. & Succ. Journ. Amer. 36. 163., 1964

Cárdenas, Cactus and Succulent Journal [U. S. ] 28: 112, f. 75-76, 1956

Salm-Dyck, Cact. Hort. Dyck. ed. I. 48, nomen.,

C. Gay, Flora Chilena 3: 29, 1847

Pfeiffer, Enumeratio Diagnostica Cactearum 144, 1837

Lemaire, Cact. Gen. Sp. Nov. 73,

D. Griffiths, Annual Report of the Missouri Botanical Garden 21: 168-169, t. 22, upper fig., 1910

K. Schumann, Monatsschrift fur Kakteenkunde 14: 26, 1904

A. Castellanos, Lilloa 23: 7, f. 2, 1950

Weing., Monatsschrift der Deutschen Kakteen-Gesellschaft, E. V., Sitz Berlin 9: 167, 1929

Rose, Smithsonian Miscellaneous Collections 50: 507, 1908

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 202, f. 250-251, 1919

(Werdermann) P.J. Braun & E. Esteves Pereira, Succulenta, 74 (3): 133: 1995

Engelmann ex Coulter, U. S. Dept. Agric. Contrib. Nat. Herb. 3. 423.,

Spegazzini, Contr. Fl. Ventana, 30., 1896

(Backeberg) Backeberg, Cactac. Handb. Kakteenk. 6. 3619, 1962

Hort. ex K. Schumann, Gesamtb. Kakt. 1. 694,

K. Schumann, Monatsschrift fur Kakteenkunde 149, 1899

Orcutt, West American Scientist 10: 81, 1896

(Engelmann) J.M. Coulter, Contributions from the U. S. National Herbarium 3: 446, 1896

Pfeiffer, Otto & Dietr. Allgemeine Gartenzeitung 6. 276., 1838

Engelmann, American Journal of Science, and Arts, ser. 2, 14 (42): 339, 1852

(Engelmann ex Parish) Borg, Cacti 107, 1952

(Engelmann) L.D. Benson, Cact. Calif. 88, 1969

A. Berger, Hort. Mortol. 234., 1912

S. Arias & Gama-Lopez, Flora del Valle de Tehuacan-Cuicatlan 14: 125, 1997

Salm-Dyck, Cact. Hort. Dyck. ed. 2. 238.,

Salm-Dyck, Hortus Dyckensis ou Catalogue des Plantes . 364, 1834

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 101-102, t. 17, f. 1, 1919

R. Philippi, Linnaea. 33. (-65) 82., 1864

J. A. Purpus, Mitteil. Deutsch. Dendrol. Ges., 63., 1925

Hort. Angl. ex Salm-Dyck, Cact. Hort. Dyck. ed. 2. 71,

Spegazzini, Nova Addenda ad Floram Patagonicam . 4: 291, 1902

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 219, f. 286, 1919

Salm-Dyck, Allgemeine Gartenzeitung 13: 387, 1845

(Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 6., 1958

(Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 6., 1958

D. Griffiths, Annual Report of the Missouri Botanical Garden 22: 33, 1912

Le Conte ex Chapman, Flora of the southern United States 145, 1860

Kuntze, Rev. Gen. 260, 1891

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 217, f. 281, 1919

Engelmann, Memoirs of the American Academy of Arts and Science 4: 52, 1849

(Engelm.) Schelle, Proceedings of the American Academy of Arts and Sciences 3: 293, 1856 [1857]

Engelmann ex W.T. Marshall, Des. Bot. Gard. Ariz. Sci. Bull. 1: 94, 1950

(Engelmann & J. M. Bigelow) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 125, 1969

(Clokey) Backeberg, Cactaceae (Backeberg) 1: 508., 1958

(Rose) H. Bravo Hollis, Cactaceas y suculentas mexicanas 19 (2): 47, 1974

Fosberg, Bull. S. Calif. Acad. Sci. 33: 102., 1934

(Griffiths) L.D. Benson & Walk., Annals of the Missouri Botanical Garden 52: 265, 1965

L.D. Benson, Cact. Succ. J. (U. S. A.), 46 (2): 79., 1974

(J.M. Coulter) L.D. Benson, The cacti of Arizona 21, 1969

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 293, 1856 [1857]

(Engelmann & J. M. Bigelow) Fosberg, Bulletin of the Southern California Academy of Sciences 33: 103, 1934

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 293, 1865

Fosberg, Bulletin of the Southern California Academy of Sciences 33: 102, 1934

(M.S. Anthony) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 125, 1969

(Griffiths) L.D. Benson, Cact. Succ. J. (U. S. A.), 46 (2): 79, 1974

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 437, 1896

(Griffiths) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 125, 1969

Haage & Sch. ex Bois, Dict. Hortic. 939,

Salm-Dyck ex Foerst., Handb. Cact. ed. I. 508, nomen.,

(L.) Mill., The Gardeners Dictionary: eighth edition, 1768

Urb., Fedde, Repert. 16. 35, 1919

Marn.-Lapost., Cactus (Paris) 15 (66): 4, 1960

Hort., ex Gartenfl. 36. 39., 1887

Parl., ex K. Schum. Gesamtb. Kakt. 741, in syn., 1898

F.A.C. Weber, Dictionaire d'Horticulture 894, 1898

Backeberg, Descr. Cact. Nov. 10., 1956

B.D. Parfitt, Rhodora 99 (899): 223-227, f. 1, 1997

Hort. ex Salm-Dyck, Cact. Hort. Dyck. ed. I. 47,

Small, The Cactaceae; descriptions and illustrations of plants of the cactus family 4: 258-259, f. 236, 1923

(Karwinski ex Pfeiffer) F.A.C. Weber, Bulletin du Museum d'Histoire Naturelle 4: 166, 1898

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 188-189, f. 232, 1919

(F. Ritter) Iliff, Cactaceae Consensus Initiatives 2: 9, 1997

Salm-Dyck, Enum. Diagn. Cact. 103,

(Salm-Dyck) Schelle, Kakteen (Schelle) 58., 1926

(Engelmann) Schelle, Kakteen (Schelle) 72., 1926

(Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 6., 1958

Salm-Dyck, Cact. Hort. Dyck. 245. 1850, 1849

Salm-Dyck, Cact. Hort. Dyck. 245. 1850, 1849

(Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 6., 1958

Griffiths, Bull. Torr. Bot. Cl., 43. 526., 1916

Rose, Smithsonian Miscellaneous Collections 50: 524, 1908

Backeberg, Cactus, Paris, No. 75, 32; Backeb. Cactac. Handb. Kakteenk. 6. 3615 (1962). 1962

Otto & Pfeiffer, Enumeratio Diagnostica Cactearum 174, 1837

Britton & Rose, Smithsonian Miscellaneous Collections 50: 523, 1908

Haworth, Synopsis plantarum succulentarum . 82, 1819

J. M. Coulter, Contr. U. S. Natl. Herb. 3: 437., 1896

(Engelmann) B.D. Parfitt, Cactus and Succulent Journal [U. S. ] 70 (4): 188, 1998

DJF 1223 

J. M. Coulter, Contr. U. S. Natl. Herb. 3: 436., 1896

(Engelmann & J.M. Bigelow) B.D. Parfitt, Cact. Succ. J. (U. S. A.), 70 (4): 188: 1998

J. A. Purpus, Mitt. Deutsch. Dendrol. Ges.: 63. 1925, 1925

(Engelmann & J.M. Bigelow) B.D. Parfitt, Cact. Succ. J. (U. S. A.), 70 (4): 188: 1998

(Britton & Rose) L.D. Benson, The cacti of Arizona 20, 1969

(L.D. Benson) B.D. Parfitt, Cact. Succ. J. (U. S. A.), 70 (4): 188: 1998

J. M. Coulter, Contr. U. S. Natl. Herb. 3: 436., 1896

(Engelmann & J. M. Bigelow) L.D. Benson, The cacti of Arizona 20, 1969

J. A. Purpus, Mitt. Deutsch. Dendrol. Ges.: 63. 1925, 1925

(K. Schumann) Backeberg, Die Cactaceae 1: 607, 1958

J. A. Purpus, Mitt. Deutsch. Dendrol. Ges.: 63. 1925, 1925

J. A. Purpus, Mitt. Deutsch. Dendrol. Ges.: 62. 1925, 1925

(Engelmann & J. M. Bigelow) J.M. Coulter, Contributions from the U. S. National Herbarium 3: 437, 1896

J.M. Coulter, Contr. U. S. Natl. Herb. 3: 437., 1896

Haworth, Syn. Pl. Succ. 190.,

Small, Manual of the Southeastern Flora 906, 1506, 1933

Hort. Angl. ex Pfeiffer, Enum. Diagn. Cact. 103,

Brandegee, Dictionaire d'Horticulture 899, 1898

Cárdenas, Lilloa 23: 25, f. 4, 1950

Salm-Dyck, Cacteae in Horto Dyckensi Cultae [ed. 1849] 236, 1849[1850]

Forbes, Hort. Tour Germ. 160, nomen.,

Spegazzini, Anales de Sociedad Cientifica Argentina 99: 103, 1925

Sabine ex Pfeiffer, Enum. Diagn. Cact. 155,

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 292, 1856 [1857]

Engelmann, American Journal of Science, and Arts, ser. 2, 14 (42): 338, 1852

Monville & Lemaire ex Forster, Handbuch der Kakteenkunde 478, 1846

Lemaire, Handb. Cact. ed. I. 508, nomen.,

Hort. ex Salm-Dyck, Cact. Hort. Dyck. ed. 2. 67,

Salm-Dyck, Obs. Bot. 7., 1822

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

Pfeiffer, Enumeratio Diagnostica Cactearum 156, 1837

J.C. Wendl. ex Pfeiffer, Enumeratio Diagnostica Cactearum 149, 1837

A. Castellanos, Physis, 9. 101., 1928

Engelmann, Trans. St. Louis Acad. 2: 201, 1863

Pfeiffer, Otto & Dietr. Allgemeine Gartenzeitung 8. 497., 1840

De Candolle, Memoires du Museum d'Histoire Naturelle 17: 119, 1828

Rose, Smithsonian Miscellaneous Collections 50: 521, 1908

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

R. Fries, Nov. Act. Soc. Sci. Upsal. Ser. 4. 1. no. 1, 123.,

(Haworth) Haworth, Synopsis plantarum succulentarum . 195, 1812

Pfeiffer 

Auct., Just, Jahresb. 24. 2. 380, sphalm.,

-> pycnantha v. margaritana 

Engelmann, Contributions from the U. S. National Herbarium 3 (7): 423-424, 1896

J.M. Coulter, Contributions from the U. S. National Herbarium 3: 424, 1896

(Wiegand & Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13 (1): 5, 1958

Rose, Contributions from the U. S. National Herbarium 12 (7): 292, t. 26, f. 35, 1909

D. Griffiths, Bulletin of the Torrey Botanical Club 43: 90, 1916

(Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 6., 1958

K. Schumann, Gesamtbeschreibung der Kakteen 694, 1898

(F. Ritter) P.J. Braun & E. Esteves Pereira, Succulenta (Netherlands) 74 (3): 133, 1995

K. Schumann, Gesamtbeschreibung der Kakteen 746, 1898

F.A.C. Weber (pro sp.), Dictionaire d'Horticulture 894, 1898

F.A.C. Weber, Dictionaire d'Horticulture 894, 1898

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 295, 1856 [1857]

(Engelmann & J. M. Bigelow) Schelle, Handbuch der Kakteenkultur. 50, 1907

(Engelmann ex J.M. Coulter) Schelle, Handbuch der Kakteenkultur. 50, 1907

(Engelmann ex Rumpler) Schelle, Handbuch der Kakteenkultur. 50, 1907

Engelmann  1856

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 295, 1856

Engelmann, Rep. Expl. Surv. Pacific Ocean, Botany 4: 55, t. 11, f. 1, 1857

Haage & Schmidt, Verz. Blumenz. 1915: 29, 1915

Engelmann, Pacif. Rail. Rep. 4. 41. t. 11.,

Späth 

Engelmann 

Engelmann 

Engelmann, Proc. Amer. Acad. Arts 3: 295., 1856

Engelmann & J. M. Bigelow 

Engelmann 

R. Philippi, Anales del Museo Nacional. Santiago de Chile 1891: 27, 1891

Engelmann, American Journal of Science, and Arts, ser. 2, 14 (42): 339, 1852

Salm-Dyck, Dyck. Bot. Gart. 360, 1837

(Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13. 6, 1958

F.A.C. Weber, Dictionaire d'Horticulture 896, 1898

(Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

K. Schumann, Monatsschrift fur Kakteenkunde 118, 1899

D. Griffiths, Monatsschrift fur Kakteenkunde 23: 131, 1913

(Griffiths) G. D. Rowley, Natl. Cact. Succ. J. 13: 25., 1958

Griffiths, Proc. Biol. Soc. Wash., 29. 12., 1916

Mackensen, Bulletin of the Torrey Botanical Club 39: 292, 1912

Wiggins & Rollins, Contributions from the Dudley Herbarium 3 (8): 275-277, t. 59, f. 1-2, 1943

Espinosa, Bol. Mus. Nac. Chile, 18. 31., 1940

Bello, Anales de la Sociedad Espanola de Historia Natural 10: 277, 1881

Karwinski ex W. Baxt., Loud. Hort. Brit. Suppl. 3. 599, nomen.,

Spegazzini, Anal. Mus. Buenos Aires, 11. 517., 1905

Lemaire, Cact. Aliq. Nov. Desc. 35,

K. Schumann, La Semaine Hort., 1897

Boissev. & C. Davidson, Cactus and Succulent Journal [U. S. ] 15: 138, 1943

Rehder, J. Arnold Arbor. 7: 149., 1926

J.G. Ortega, Fl. Ind. Sinaloa, 1929

Hosseus, Kaktus-ABC, 429, indice, 113, in syn., 1935

Griffiths, Proc. Biol. Soc. Wash., 27. 26., 1914

A. Berger, Kakteen 68, 1929

Backeberg, Die Cactaceae 6: 3628, 1962

J.G. Ortega, Fl. Indig. Sinaloa, 1929

Cárdenas, Cactus and Succulent Journal [U. S. ] 42: 32, 1970

Wendland, Pfeiffer, Enum. Cact. 165, 1837

Schelle, Handb. Kakteenkult. [Schelle] 57., 1907

(Griffiths) Sanchez-Mejorada, Cactaceas y suculentas mexicanas 17 (4): 119, 1972

(F.A.C. Weber) H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 119, 1972

Backeberg, Descr. Cact. Nov. 10., 1956

Schelle, Handb. Kakteenkult. [Schelle] 57., 1907

(Rose) K. Hammer, Kulterpflanze, 24: 266, 1976

G.E. Lindsay, Cactus and Succulent Journal [U. S. ] 14: 56, 1942

Schumann, Ges. Kakt. 764, 1898, 1828

Mackensen, Bulletin of the Torrey Botanical Club 38: 142, 1911

K. Schumann, Gesamtb. Kakt. 1. 686,

(Heinrich & Backeberg) D.R. Hunt, Cactaceae Consensus Initiatives 4: 6, 1997

K. Schumann, Gesamtbeschreibung der Kakteen 652, 1899

Voigt, Hort. Suburb. Calc. 62,

Salm-Dyck ex DC., Prodromus Systematis Naturalis Regni Vegetabilis 3: 474, 1828

Davidson, Bulletin of the Southern California Academy of Sciences 15 (2): 33, f. s. n. [pg. 32], 1916

Engelmann, U. S. Dept. Agric. Contrib. Nat. Herb. 3. 424.,

Salm-Dyck ex Foerst., Handb. Cact. ed. I. 493,

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 298, 1856 [1857]

M. S. Anthony, American Midland Naturalist 55: 240, 1956

D. Griffiths, Proc. Biol. Soc. Wash., 27. 26., 1914

A. Castellanos, Lilloa 9: 213, 1943

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 94-5, f. 111, 1919

Hort. ex A. Berger, Hort. Mortol. 235., 1912

Nutt., A Flora of North America: containing . 1 (4): 555, 1840

Scheinvar, Anales del Instituto de Biologia de la Universidad Nacional Autonoma de Mexico 47-53: 123, f. 1-7, 1984

Pfeiffer, Enumeratio Diagnostica Cactearum 147, 1837

A. Castellanos, Lilloa 27: 85, f. 2, 1955

Sprengel ex Pfeiffer, Enum. Diagn. Cact. 141,

Parm. ex Pfeiffer, Enumeratio Diagnostica Cactearum 172, 1837

(K. Schumann) G. D. Rowley, Natl. Cact. Succ. J. 13: 25., 1958

(F. A. C. Weber) G. D. Rowley, Natl. Cact. Succ. J. 13: 25., 1958

Hort. ex Forbes, Hort. Tour Germ. 158, nomen.,

Britton & Standley, Journal of the Washington Academy of Sciences 15 (5): 104, 1925

D.R. Hunt, Cactaceae Consensus Initiatives 4: 6, 1997

J.P. Rebman, Haseltonia, 6: 17 ( publ. 1999), 1998

C. Z. Nelson, Gatesburg Reg., 1915

D. Griffiths, Proceedings of the Biological Society of Washington 27: 26, 1914

(E. M. Baxter) Helia-H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 119, 1972

(Baxter) H. Bravo Hollis, Cact. Suc. Mex. 17 (4): 119, as 'santamaria', 1972

(Griffiths & Hare) Rose, Smithsonian Miscellaneous Collections 52: 195, 1908

Griffiths ex L. Scheinvar, Phytologia, 49 (4): 328, 1981

(F. Ritter) P.J. Braun & E. Esteves Pereira, Cactus and Succulent Journal [U. S. ] 63 (2): 82, 1991

P.J. Braun & E. Esteves Pereira, Cactus and Succulent Journal [U. S. ] 67 (2): 108-112, f. 1-7, 1995

P.J. Braun & E. Esteves Pereira, Cact. Succ. J. (U. S. A.), 63 (6): 311, 1991

P.J. Braun & E. Esteves Pereira, Cact. Succ. J. (U. S. A.), 63 (3): 124, 1991

J.T. Howell, Proceedings of the California Academy of Sciences, Series 4, 21: 45, 1933

F.A.C. Weber, Dictionaire d'Horticulture 895, 1898

F.A.C. Weber, Dictionaire d'Horticulture 898, 1898

Hort. Berol. ex Salm-Dyck, Cact. Hort. Dyck. ed. I. 48, nomen.,

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 304, 1856 [1857]

(Engelmann) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 124, 1969

A. Castellanos & H.V. Lelong, Lilloa 10: 400, 1944

Spegazzini, Anales del Museo Nacional de Buenos Aires 11: 511, 1905

K. Schumann, Monatsschrift fur Kakteenkunde 9: 148, 1899

(Backeberg) Moran, Gentes Herbarum; occasional papers on the kind of plants 8 (4): 326, 1953

(Lemaire) L. Benson, Cact. Succ. J. (U. S. A.), 46 (2): 80, 1974

Engelmann, Boston Journal of Natural History 6 (2): 208, 1850

(Engelmann) Moran, Baileya. 19 (4): 166, 1975

(Lemaire) Moran, Baileya. 19 (4): 166, 1975

Borg, Cacti 67, 1937

F. Phil., Botanische Zeitung. Berlin 26: 861, 1868

C. Z. Nelson, Transactions of the Illinois State Academy of Science 12: 120, 1919

Griffiths, Bull. Torr. Bot. Cl., 43. 89., 1916

Roezl, La Belgique Horticole 24: 39, 1874

Engelmann, Boston Journal of Natural History 6 (2): 206, 1850

Britton & Rose 

L. Scheinvar, An. Inst. Biol. Univ. Nac. Auton. Mex., Bot., 45 (1): 85 ( publ. 1975). 1974

(Backeberg) H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 118, 1972

(S.D.) Rowley, Rep. Pl. Succ., 22 (Regnum Veg., 87): 10, without basionym page, 1973

H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 119, 1972

Britton & Rose 

G. Don ex Salm-Dyck, Hort. Dyck. 363.,

Salm-Dyck, Hort. Dyck. 363., 1834

Salm-Dyck, Cact. Hort. Dyck. 67, 237. 1850, 1849

Engelmann, American Journal of Science, and Arts, ser. 2, 14 (42): 338, 1852

(Backeberg) W. T. Marsh., Cact. & Succ. Journ. Amer. 17. 114, 1945

Engelmann, Cacteae in Horto Dyckensi Cultae [ed. 1849] 239, 1849[1850]

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 72, 1919

(Backeberg) G.D. Rowley, Cact. Succ. J. Gr. Brit., 44 (1): 2: 1982

A. Nelson, Gatesburg Reg., 5680

Backeberg, Cactus Journ. 1. 7., 1932

D. Griffiths, Annual Report of the Missouri Botanical Garden 21: 173-174, t. 28, 1910

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 96-97, f. 114, 1919

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 221, f. 294, 1919

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 134-135, f. 169, 1919

(Otto) F.A.C. Weber, Bulletin du Museum d'Histoire Naturelle 4: 165, 1898

Steud., Nomenclator Botanicus. Editio secunda 2: 222, 1841

(Daston) Rowley, Nat. Cact. & Succ. Journ. 13 (1): 5, 1958

F.A.C. Weber, Dictionaire d'Horticulture 898, 1898

Graessn., Hauptverz., 10, nomen., 1937

C. F. Forster, Hamburger Garten- und Blumenzeitung 17: 167, 1861

(K. Schumann) G.D. Rowley, National Cactus and Succulent Journal 13 (2): 25, 1958

(Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (2): 25, 1958

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 300, 1856 [1857]

Engelmann, Transactions of the Academy of Science of St. Louis 11: 199, 1863

Rafin., Atl. Journ. 147 (Quid ?).

Karwinski ex Salm-Dyck, Cact. Hort. Dyck. ed. I. 46,

R. Philippi, Linnaea 30: 211, 1859

M.S. Anthony, American Midland Naturalist 55: 246, f. 22-23, 1956

(Engelmann) Toumey, Botanical Gazette 25: 119, 1898

(Haworth) DC. 

Toumey, Botanical Gazette 25: 119, 1898

Miller, The Gardeners Dictionary: eighth edition Opuntia no. 88, 1768

D. Griffiths, Proceedings of the Biological Society of Washington 27: 24, 1914

Salm-Dyck, Hortus Dyckensis ou Catalogue des Plantes . 364, 1834

Salm-Dyck, Cact. Hort. Dyck. ed. I. 46, nomen.,

Engelmann & Bigel. ex Haage, Verz. Cact. 30.,

Hort. Angl. ex Pfeiffer, Enum. Diagn. Cact. 159,

J.G. Ortega, Fl. Ind. Sinaloa 6, 1929

Griffiths, Bull. Torr. Bot. Cl., 43. 91., 1916

Bullock, Cactus Journal [London] 8: 15, 1939

Engelmann in Emory, Notes Milit. Reconn. 2: 158., 1848

Engelmann, Notes of a Military Reconnoissance 2: 158, 1848

(Rose) L.D. Benson, Proc. Calif. Acad. Sci. 4th ser. 25: 248., 1944

(Orcutt) L.D. Benson, Cacti Ariz. 2nd. ed. 39., 1950

L.D. Benson, The cacti of Arizona 20, 64, 1969

(E. M. Baxter) L. D. Benson, Proceedings of the California Academy of Sciences, Series 4, 25: 248, 1944

De Candolle, Memoires du Museum d'Histoire Naturelle 17: 117, 1828

(Backeberg) G.D. Rowley, Nat. Cact. & Succ. Journ. 13. 4, 1958

Salm-Dyck, Cact. Hort. Dyck. ed. I. 48; ed. 2. 73,

K. Schumann, Monatsschrift fur Kakteenkunde 149, 1899

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 296, 1856 [1857]

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 289, 1856

H. Bravo Hollis, Cactaceas y suculentas mexicanas 19 (1): 27, 1974

(Coulter) K. Schumann, Gesamtb. Kakt. 679, 830, 1898

Sencke ex Haage, Verz. Cact. 30.,

Lemaire, Cactearum Genera Nova Speciesque Novae 62, 1839

(Griff.) K. Hammer, Kulterpflanze, 24: 262, 1976

(Haworth) Haworth, Synopsis plantarum succulentarum . 191, 1812

(Ker Gawl.) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 126, 1969

(Haworth) Haworth, Cactus and Succulent Journal (Los Angeles) 41 126., 1969

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 290, 1856 [1857]

(Griffiths) L.D. Benson, Cact. Succ. J. (U. S. A.), 46 (1): 79, 1974

A. Berger, Kakteen 53, la|4m. 9, 1929

D. Griffiths, Illustrated Studies in the Genus Opuntia. -- 2. 94, t. 2, f. 1, t. 11, 13, f. 4, 1909

Schott, Enumeratio Diagnostica Cactearum 167, 1837

(F.M. Knuth) H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 118, 1972

Engelmann, Proceedings of the American Academy of Arts and Sciences 3 302., 1857

(F. Reichenbach & K. Schumann) H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 118, 1972

(Salm-Dyck) Z.Y. Li, Guihaia, 1 (4): 42, without basionym ref.: 1981

Mill., Proceedings of the American Academy of Arts and Sciences 3 289., 1857

Engler 

Link ex Steud., Nom. ed. 2. 2. 222, nomen.,

Spegazzini, Anales de Sociedad Cientifica Argentina 99: 104, 1925

R. Fries, Nova Acta Regiae Societatis Scientiarum Upsaliensis, ser. 4, 1 (1): 122, 1905

(Muehlenpf.) Engelmann, Gardener's Chronicle & Agricultural Gazette 19: 627, 1883

(Berg.) Rowley, Repert. Pl. Succ. (I. O. S.), 23: 8 ( publ. 1974), without basionym date, 1972

G. Don, Loudon's Hortus Britannicus. Second edition, 196, 1830

(F. Ritter) P.J. Braun & E. Esteves Pereira, Succulenta, 74 (3): 133: 1995

(F. Ritter) P.J. Braun & E. Esteves Pereira, Succulenta, 74 (3): 133: 1995

(R.E. Fries) Backeberg 

Salm-Dyck, Cact. Hort. Dyck. 67. 1850, 1849

Monville ex Lemaire, Cact. Gen. Sp. Nov. 71., 1839

Monville ex Lemaire, Cact. Gen. Sp. Nov. 71., 1839

(Spegazzini) Backeberg, Cactaceae (Backeberg) 1: 414., 1958

D. Griffiths, Proceedings of the Biological Society of Washington 29: 13, 1916

Pfeiffer ex Foerst., Handb. Cact. ed. I. 470,

Engelmann, Contributions from the U. S. National Herbarium 3: 423, 1896

R. Philippi, Anales del Museo Nacional. Santiago de Chile 1: 27, 1891

D. Griffiths, Annual Report of the Missouri Botanical Garden 22: 34, t. 11, 15, 1912

Cárdenas, Lilloa 23: 18, 1950

Britton & Rose, Smithsonian Miscellaneous Collections 50: 520, 1908

S. Arias & L.U. Guzman, Flora del Valle de Tehuacan-Cuicatlan 14: 131, 1997

(H. Bravo Hollis) H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 118, 1972

Auct., Just, Jahresb. 24. 2. 380, sphalm.,

Small, Manual of the Southeastern Flora 908, 1933

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 294, 1856

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 4, 1958

F. Cels ex F.A.C. Weber, Dictionaire d'Horticulture 898, 1898

Engelmann ex J.M. Coulter, Contributions from the U. S. National Herbarium 3: 448, 1896

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 309, 1856

K. Schumann, Monatsschr. Kakteenk. 8: 70, pl., 1898

F. A. C. Weber, Bull. Mus. Hist. Nat. (Paris) 10: 389., 1904

F.A.C. Weber 

Toumey, Gard. & Forest 9: 432., 1896

D. Griffiths, Illustrated Studies in the Genus Opuntia. -- 2. 92-93, t. 9, 13, f. 1, 1909

Thornber & Bonker, The Fantastic Clan 133, 148, f. p. 135, 1932

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 308, 1856

(Engelmann) Felger & Lowe, J. Arizona Acad. Sci. 6: 83., 1970

(Britton & Rose) H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 119, 1972

(F. Ritter) D.R. Hunt, Cactaceae Consensus Initiatives 4: 6, 1997

J. M. Bigelow  1856

Bigel., in. Pacif. Rail. Rep. 4. 11,

Backeberg, Backeb. & F. M. Knuth, Kaktus-ABC, 128, 411., 1935

Backeberg, Cactaceae (Backeberg) 6: 3617., 1962

A. Berger, Monatsschrift fur Kakteenkunde 20: 81, 1910

Salm-Dyck, Observationes Botanicae in Horto Dyckensi 3: 8, 1822

(DC.) H. Bravo Hollis, Cactaceas y suculentas mexicanas 20 (4): 96, 1975

L. Scheinvar, Phytologia, 49 (4): 332, 1981

(J. G. Ortega) Backeberg, Cactaceae (Backeberg) 1: 542., 1958

(J. G. Ortega) Backeberg, Cactaceae (Backeberg) 1: 542. as Spranguei, 1958

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 293, 1856 [1857]

(Engelmann & J. M. Bigelow) Backeberg, Die Cactaceae 1: 483, 1958

Rose, Contributions from the U. S. National Herbarium 12: 402, 1909

Britton, Torreya 11: 152, 1911

Coulter, U. S. Dept. Agric. Contrib. Nat. Herb. 3. 434.,

Griffiths & Hare, Bull. Agric. Exp. Sta. New Mex. Coll. Agric. Mech. Arts 60: 81., 1906

(Willd.) Sweet, Hortus Britannicus 172, 1826

D. Griff., Monatsschr. Kakteenk., 23. 137., 1913

(Engelmann & J. M. Bigelow) Britton & Rose, Smithsonian Miscellaneous Collections 50: 535, 1908

D. Griffiths, Illustrated Studies in the Genus Opuntia. -- 2. 85-86, t. 4, upper f., 1909

Haworth, Suppl. Pl. Succ. 80.,

(Wiegand & Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13 (1): 5, 1958

R. Philippi, Anal. Mus. nac. Chile 28., 1891

Pfeiffer, Enumeratio Diagnostica Cactearum 146, 1837

Pfeiffer, Enum. Diagn. Cact. 146., 1837

(F. Ritter) D.R. Hunt, Cactaceae Consensus Initiatives 4: 6, 1997

(L.) Mill., The Gardeners Dictionary: eighth edition Opuntia no. 3, 1768

Gentry, Allan Hancock Atlantic Exped., Rep. 2, Carrib. Mar. Alg. Rep. no. 6: 11., 1948

Orcutt, W. Amer. Sci. 7: 157., 1891

Spegazzini, An. Soc. Ci. Argent. 99. 107., 1925

(Lehmann) Link & Otto, Enumeratio Diagnostica Cactearum 170, 1837

(F. Ritter) A.E. Hoffmann J., Cactaceas en la flora silvestre de Chile: 242: 1989

(Engelmann & J. M. Bigelow) L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 124, 1969

Salm-Dyck, Cact. Hort. Dyck. 73, 250. 1850, 1849

Gibbes, Proceedings of the Elliott Society of Natural History 1: 272, 1859

Britton & Shafer ex Bold., The Flora of the Dutch West Indian Islands 2: 81, 1914

Small, Manual of the Southeastern Flora 910, 1933

Small, The Cactaceae; descriptions and illustrations of plants of the cactus family 4: 265, 1923

Lemaire, Cact. Aliq. Nov. Desc. 36.,

Salm-Dyck, Cact. Hort. Dyck. 71. 1850, 1849

Hort. ex Schelle, Handb. Kakteenk. 58, in syn., 1907

Weing., Kakteenkunde 1933: 71, 1933

Engelmann x O. kelvinensis 

Steud., Nom. ed. 2. 2. 222,

D. Griffiths, Monatsschrift fur Kakteenkunde 23: 139, 1913

D. Griffiths, Monatsschrift fur Kakteenkunde 22: 32, 1912

Backeberg, Blatter fur Kakteenforschung 1937-7, gen. 10, sp. 7, 1937

Werdermann, Repertorium Specierum Novarum Regni Vegetabilis 29: 229, 1931

F.A.C. Weber, Dictionaire d'Horticulture 2: 896, 1898

J.A. Purpus, Monatsschrift fur Kakteenkunde 19: 133, 1909

Spegazzini, Anal. Mus. Buenos Aires, 11. 516.,

Engelmann, Memoir of a Tour to Northern Mexico 100, 1848

D. Griffiths, Proceedings of the Biological Society of Washington 27: 24, 1914

(J.M. Coulter) Britton & Rose (pro sp.), Smithsonian Miscellaneous Collections 50: 532, 1908

Parish in Jeps., Man. Fl. Pl. Calif. 657., 1925

F.A.C. Weber, Bulletin du Museum d'Histoire Naturelle 10: 389, 1904

(Griffiths) H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 119, 1972

(Rose) H. Bravo Hollis, Cactaceas y suculentas mexicanas 17 (4): 119, 1972

Hort ex Cels, Cat. Cactees, 19, nomen., 1858

F. Cels ex F.A.C. Weber, Dictionaire d'Horticulture 898, 1898

F. A. C. Weber in Bois, Dict. Hort. 2: 898., 1898

Borg in Borg, Cacti (Borg) 62., 1937

(Spegazzini) G. D. Rowley, Natl. Cact. Succ. J. 13: 25., 1958

(Backeberg) G.D. Rowley, Natl. Cact. Succ. J. 13: 4., 1958

Engelmann ex J.M. Coulter, Contributions from the U. S. National Herbarium 3: 452, 1896

Weing., Kakteenkunde 1933: 72, 1933

Salm-Dyck, Allgemeine Gartenzeitung 13: 388, 1845

(Backeberg) Krainz in Krainz, Kakteen (H. Krainz) 36: C 8. B., 1967

(Backeberg) Krainz in Krainz, Kakteen (H. Krainz) 36: C 8. B., 1967

(Cárdenas) G. D. Rowley, Natl. Cact. Succ. J. 13: 25., 1958

D. Griffiths, Annual Report of the Missouri Botanical Garden 22: 28, 1912

Rose, Contributions from the U. S. National Herbarium 12 (7): 293, t. 27, t. 36, 1909

Engelmann, Emory, Notes Mil. Reconnois. 157.,

(Rose) Lodé, Fichier Encycl. Cact. Autres Succ., cf. Repert. Pl. Succ. (I. O. S.), Sér. 17: no. (1996), without basionym ref., species name not validly publ.; 47: 11 (publ. 1997): 1996

L.D. Benson, Cactus and Succulent Journal [U. S. ] 41: 125, 1969

(F.A.C. Weber) L.D. Benson, The cacti of Arizona 21, 1969

(Engelmann) L.D. Benson, The cacti of Arizona 21, 1969

(Griffiths & Hare) L.D. Benson, The cacti of Arizona 21, 1969

Tenore, Atti 7 Adun. Sci. Ital. 886, 1846

Link & Otto, Handbuch der Kakteenkunde 990, 1846

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 55, f. 65, 1919

(F. Ritter) P.J. Braun & E. Esteves Pereira, Succulenta (Netherlands) 74 (3): 133, 1995

(F. Ritter) P.J. Braun & E. Esteves Pereira, Succulenta, 74 (3): 133: 1995

(F. Ritter) P.J. Braun & E. Esteves Pereira, Succulenta (Netherlands) 74 (3): 133, 1995

(F. Ritter) P.J. Braun & E. Esteves Pereira, Succulenta, 74 (3): 133: 1995

Rose (pro sp.), Smithsonian Miscellaneous Collections 52: 153, 1908

Miller, The Gardeners Dictionary: eighth edition Opuntia no. 1, 1768

F. A. C. Weber in Bois, Dict. Hort. 2: 894., 1898

(Cons ex F. A. C. Weber) Backeberg, Cactaceae (Backeberg) 1: 400., 1958

Salm-Dyck, Observationes Botanicae in Horto Dyckensi 3: 9, 1822

Salm-Dyck, Observationes Botanicae in Horto Dyckensi 3: 9, 1822

Schelle, Handbuch der Kakteenkultur. 50, 1907

(Engelm.) A. Gray, A Manual of the Botany of the Northern United States. Second Edition 136, 1856

A. Web., Bois, Dict. Hortic. 895.,

(F. Ritter) D.R. Hunt, Cactaceae Consensus Initiatives 4: 6, 1997

A. Weber, Bull. Mus. Hist. Nat. Par., 393., 1904

Spegazzini, Anales del Museo Nacional de Buenos Aires 11: 509, 1905

(Backeberg) G. D. Rowley, Natl. Cact. Succ. J. 13: 25., 1958

Backeberg, Blatter fur Kakteenforschung 1935-1938, gen. 10, subg. 4-a-b, sp. 2, 1935

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 116, 1919

U. Eggli, Bradleya; Yearbook of the British Cactus and Succulent Society 10: 90, 1992

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 4: 255-256, f. 234, 1923

Engelmann & J. M. Bigelow, Proceedings of the American Academy of Arts and Sciences 3: 307, 1856 [1857]

Peebles, Journal of the Washington Academy of Sciences 30 (11): 473, 1940

Engelmann, Proc. Amer. Acad. Arts 3: 307., 1856

(Clokey) L.D. Benson, The cacti of Arizona 20, 1969

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 307, 1856 [1857]

(Britt. & Rose) L. Benson, Cact. Succ. J. (U. S. A.). 46 (2): 79, 1974

E.M. Baxter, Calif. Cactus 37, 1935

E.M. Baxter, Calif. Cactus 39, 1935

(Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13 (1): 5, 1958

L.D. Benson, The cacti of Arizona 19, 32, 1969

Britton & Rose, The Cactaceae; descriptions and illustrations of plants of the cactus family 1: 172, f. 211, 1919

(Backeberg) Rowley, Nat. Cact. & Succ. Journ. 13. 6, 1958

A. Berger, Botanische Jahrbucher fur Systematik, Pflanzengeschichte und Pflanzengeographie 36: 455, 1906

(L.D. Benson) M.S. Baker, Madrono; Journal of the California Botanical Society 39: 108, 1992

Backeberg, Descriptiones Cactacearum Novarum 10, 1956

D. Griffiths, Annual Report of the Missouri Botanical Garden 21: 171-172, t. 26, upper fig., t. 27, 1910

(E. M. Baxter) Peebles, Desert Plant Life 9: 45, 1937

Engelmann, Proceedings of the American Academy of Arts and Sciences 3: 308, 1856 [1857]

D. Griffiths, Annual Report of the Missouri Botanical Garden 21: 166-167, t. 20, lower fig., 1910

K. Schumann, Gesamtb. Kakt. 1. 735.,

D. Griffiths, Proceedings of the Biological Society of Washington 29: 15, 1916

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13 (1): 5, 1958

C. Z. Nelson, Trans. Illinois State Acad. Sci. 12: 119. Galesburg Rep. Reg. July 20, 1915, 1919

J.T. Howell, Proceedings of the California Academy of Sciences, Series 4, 21: 48, 1933

A. Berger, Hortus Mortolensis, 236, 413., 1912

Scheinvar, Cact. Suc. Mex. 44 (4): 88 (-93; figs. 1-6). 1999

Small, Journal of the New York Botanical Garden 20: 35, t. 226, 1919

(Rauh & Backeberg) G.D. Rowley, National Cactus and Succulent Journal 13: 6, 1958

D. Griffiths, Bulletin of the Torrey Botanical Club 43: 86, 1916